1. of New Mexico Museum of Southwestern Biology (MSB 72651).
New county record (Degenhardt et al. 1996. Amphibians and Herpetological Review, 2010, 41(4), 520–523.
© 2010 by Society for the Study of Amphibians and Reptiles
Reptiles of New Mexico. Univ. New Mexico Press, Albuquerque.
431 pp.). The specimen was found under a small board on moist Noteworthy Distribution Records of Reptiles
soil. Other reptile species noted nearby included Diadophis punc- from Western Panamá
tatus, Opheodrys vernalis, Sceloporus poinsettii, S. cowlesi, and
Phrynosoma hernandesi. During September 2005 EM found and
SEBASTIAN LOTZKAT*1, 2
photographed an adult female T. lineatum at this location, but it
ANDREAS HERTZ1, 2
was not collected. LEONHARD STADLER1, 3
Submitted by IAN W. MURRAY, MSC03 2020, Department NADIM HAMAD1, 2
of Biology, 1 University of New Mexico, Albuquerque, New ARCADIO CARRIZO4
Mexico 87131-0001, USA (e-mail: imurray@unm.edu); ERIK and
GUNTHER KÖHLER1
McCORMICK, 1011 S Yates St., Artesia, New Mexico 88210-
2747, USA (e-mail: nmherper@hotmail.com); and CHRISTO-
1
Senckenberg Forschungsinstitut und Naturmuseum Frankfurt
Senckenberganlage 25, 60325 Frankfurt am Main, Germany
PHER NEWSOM, 5519 Amistad Rd. NE, Albuquerque, New
Mexico 87111-1549, USA (e-mail: herpsofnm@comcast.net).
2
Johann Wolfgang Goethe-University
Institute for Ecology, Evolution and Diversity, BioCampus Westend
Siesmayerstr. 70, 60323 Frankfurt am Main, Germany
TROPIDODIPSAS FASCIATA (Banded Snail Sucker). MÉXI- 3
Justus-Liebig-University, Department of Animal Ecology,
CO: GUERRERO: MuniciPalitY of ziHuataneJo: 27 km SW Val- Heinrich-Buff-Ring 26-32 (IFZ), 35392 Giessen, Germany
lecitos de Zaragoza, on México Hwy 134 (17.782°N, 101.478°W; 4
Universidad Autónoma de Chiriquí, Davíd, Panamá
WGS84), 333 m elev. 02 September 2008. William H. Mertz. Ver-
ified by Coleman M. Sheehy III. UTA digital collection (UTADC *Corresponding author; e-mail: lotzkat@yahoo.com
3702). New municipality record, which extends the range of the
species ca. 200 km W of its closest known localities in central Despite of its small size, Panamá is home to 251 species of rep-
Guerrero (Kofron 1987. J. Herpetol. 21:210–225). The snake was tiles, including 89 lizards and 143 snakes (Jaramillo et al. 2010;
found AOR at night in tropical deciduous forest. Köhler 2010; Köhler et al. 2010). The descriptions of new rep-
Submitted by WILLIAM H. MERTZ, Avenida La Playa, tiles continue at a steady pace (e.g., Cadle and Myers 2003; Hule-
Troncones, La Union, Guerrero, México 39270 (e-mail: Crota- bak et al. 2007; Köhler 2010; Köhler et al. 2010, 2007; Köhler
lus_mx@yahoo.com); JACOBO REYES-VELASCO, Centro and Sunyer 2008; Myers 2003; Poe and Ibañez 2007; Poe et al.
Universitario de Ciencias Biológicas y Agropecuarias, Carretera 2009; Savage and Watling 2008; Savage et al. 2008), indicating
a Nogales Km. 15.5. Las Agujas, Nextipac, Zapopan, Jalisco, that our knowledge of the Panamanian herpetofauna is still frag-
México (e-mail: jackobz@gmail.com); and CHRISTOPH I. mentary and investigative work remains to be done. This applies
GRÜNWALD, Casa Mexico Real Estate, Ajijic Plaza #1, Ajijic, to taxonomic issues as well as to biogeography. Additionally,
Jalisco, México (e-mail: cgruenwald@switaki.com). the distributions of many species occurring within Panamá are
poorly documented. Apart from information scattered among lo-
TROPIDOPHIS GREENWAYI GREENWAYI (Big Ambergris cal checklists and publications on single species, only four com-
Cay Dwarf Boa). TURKS AND CAICOS ISLANDS: Big aMBer- prehensive works exist for the country. Pérez-Santos (1999) pre-
griS caY: N end of island (21.317°N, 71.630°W; WGS84), elev. sented distribution maps with precise collecting localities for 129
3 m. 20 March 2009. J. Burgess, G. Gerber, and G. Reynolds. snake species. Köhler (2008) provided distribution maps for 238
Verified by J. Iverson. Color photographic voucher, Austin Peay reptile species occurring in Panamá. Some of the maps pinpoint
State University (APSU 19018). First published record for Big actual collection sites, but most reflected areas of generalized
Ambergris Cay, the type locality, since 1936 and only the third in- distribution. The only work considering distribution of the her-
dividual ever encountered on the island. Because it had not been petofauna among Panamá’s political subdivisions was by Young
found since 1936, despite a great deal of herpetological investiga- et al. (1999), therein summarizing the distribution of 228 spe-
tions, the species was thought to have been extirpated from the cies at the province level. However, their distributional records
Island (Iverson 1986. Carib. J. Sci. 22:191–198). The male snake were not supported by reference to voucher material, and they
was encountered at night, during a light rain, on a dirt road at the did not include the results of two extensive regional inventories
base of a rocky hillside. by Martínez and Rodriguez (1992) and Martínez et al. (1994).
Submitted by R. GRAHAM REYNOLDS, Department of Recently, Jaramillo et al. (2010) summarized the distribution of
Ecology and Evolutionary Biology, University of Tennessee, the Panamanian herpetofauna within physiographic regions and
Knoxville, Tennessee 37996, USA (e-mail: rgraham@utk.edu); vegetation zones.
GLENN P. GERBER, San Diego Zoo’s Institute for Conser- During field work on reptiles conducted in western Panamá
vation Research, Escondido, California 92027, USA (e-mail: between May and August 2008, we visited localities within the
ggerber@sandiegozoo.org); and JOESEPH BURGESS, Guana provinces of Bocas del Toro, Chiriquí, Veraguas, and Comarca
Tolomato Matanzas National Estuarine Research Reserve, Ponte Ngöbe-Buglé (a recently established indigenous autonomous re-
Vedra, Florida 32082, USA (e-mail: Joseph.Burgess@dep.state. gion that comprises areas formerly belonging to the other three
fl.us). provinces) (Fig. 1). We collected 217 samples representing 72
species and 17 of those constitute extensions of their known dis-
tributions in Panamá. We herein report distribution extensions of
520 Herpetological Review 41(4), 2010

2. (8.5095°N, 81.1173°W), 900 m elev. 2 August 2008. Leonhard
Stadler and Nadim Hamad. SMF 89577. This record extends
by 200 m, the upper confirmed elevational limit for the species
previously reported by Savage (2002) and Köhler (2008). The
presence of L. southi around Alto de Piedra was documented by
Martínez and Rodriguez (1992). However, elevational distribu-
tion for their individual records was unspecified; only a general
elevational range of between 800 m and 1450 m was given.
Hoplocercidae
Enyalioides heterolepis (Bocourt’s Dwarf Iguana). VERAGUAS:
Cerro Negro, ca. 6 km NNW of Santa Fé (8.5690°N, 81.0989°W),
700 m elev. 30 July 2008. Sebastian Lotzkat. SMF 89558. Cerro
Negro, 6 km NNW of Santa Fé (8.5693°N, 81.1022°W), 713 m
fig. 1. Generalized localities in western Panamá visited by the authors elev. 28 June 2008. Arcadio Carrizo. SMF 89955–56. First re-
in 2008: 1) Jurutungo, Chiriquí; 2) Reserva Forestal La Fortuna, Chiriquí; cords for Veraguas, extending the range more than 50 km W
3) road to Cerro Colorado, Comarca Ngöbe-Buglé; 4) La Nevera,
of the formerly westernmost record from El Copé, Coclé (Lips
Comarca Ngöbe-Buglé; 5) Cerro Mariposa (= Cerro Tute), Veraguas; 6)
Cerro Negro, Veraguas; 7) Río Chilagres, Veraguas. 1999). Köhler (2008) mentioned a record of E. heterolepis from
Bocas del Toro, which as far as we can tell is not supported by
a voucher specimen. A now lost individual from Cerro Narices
those 17 species, 11 of which are new provincial records; other
(between Cerro Negro and Río Chilagres, see Fig. 1) that was
accounts list new elevational parameters for the species.
listed as Sceloporus squamosus by Martínez et al. (1994), actu-
Abbreviations for museum collections follow those of Leviton ally represented an E. heterolepis (A. Rodriguez, pers. comm.).
et al. (1985). Elevations were rounded to the next tenth. The map
(Fig. 1) was created using DIVA-GIS and the NASA elevation Polychrotidae
datasets processed by Jarvis et al. (2006); map datum is WGS84.
Anolis auratus (Grass Anole). VERAGUAS: road between Santi-
All vouchers were verified by Javier Sunyer. Common names
ago and Santa Fé, ca. 8 km N of Santa Fé; (8.1796°N, 80.951°W),
came from Frank and Ramus (1995), except for Leposoma southi
80 m elev. 14 May 2008. Andreas Hertz. SMF 89444. First record
(Uetz 2008), Urotheca guentheri (Solórzano 2004), and by us
for Veraguas, bridging the gap between localities in Chiriquí to
herein for Anolis fortunensis and A. pseudopachypus, two species
the west and Los Santos, Colón, and Panamá to the east (Köhler
for which there are no published common names. We decided
et al. 2008; Young et al. 1999).
that for those two species, the common names should denote their
type localities; Reserva Forestal La Fortuna, Panamá, and La Ne- Anolis capito (Bighead Anole). CHIRIQUÍ: Reserva Forestal La
vera, Panamá, respectively. Fortuna (8.6744°N, 82.2161°W), 1460 m elev. 16 June 2008. An-
dreas Hertz. SMF 89449. This record increases the known eleva-
Squamata — Lizards tional limit for the species by 160 m (Köhler 2008).
Sphaerodactylidae Anolis fortunensis (La Fortuna Anole). CHIRIQUÍ: Reserva For-
Lepidoblepharis xanthostigma (Costa Rica Scaly-eyed Gecko). estal La Fortuna (8.6776°N, 82.1981°W), 1750 m elev. 27 June
VERAGUAS: Cerro Mariposa, near Alto de Piedra, ca. 3.5 km W 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89466. This
of Santa Fé (8.5161°N, 81.1185°W), 880 m elev. 12 May 2008. record extends the known elevational range of the species by 550
Andreas Hertz and Sebastian Lotzkat. SMF 89576. Cerro Negro, m (Köhler 2008).
ca. 6 km NNW of Santa Fé (8.5690°N, 81.0989°W), 700 m elev. Anolis humilis (Humble Anole). CHIRIQUÍ: Reserva Forestal
28 July 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89963. La Fortuna (8.6761°N, 82.2006°W), 1660 m elev. 27 June 2008.
Martínez and Rodriguez (1992), in their inventory of “Cerro Sebastian Lotzkat. SMF 89480. This record increases the known
Tute” (= Cerro Mariposa) and Martínez et al. (1994) in their elevational range by 60 m (Köhler 2008).
species list of the nearby mountains, Narices and La Anselma, Anolis pseudopachypus (La Nevera Anole). CHIRIQUÍ: Reser-
both reported the presence of Lepidoblepharis sp., and consid- va Forestal La Fortuna (8.6761°N, 82.2006°W), 1660 m elev.;
ered them to resemble L. xanthostigma. However, because the (8.6793°N, 82.193°W), 1800 m elev. 25 and 27 June 2008. Andre-
specific identities of these geckos were not made and the respec- as Hertz and Sebastian Lotzkat. SMF 89523, 89526, respectively.
tive voucher specimens were lost (V. Martínez pers. comm.), our First records for Chiriquí, and extends the range ca. 50 km WNW
records represent the first confirmed occurrence for the species of the type locality (La Nevera, Comarca Ngöbe-Buglé; Köhler
in Veraguas. The records also fill the gap between records to the et al. 2007). COMARCA NGÖBE-BUGLÈ: Valley north of La
west in Chiriquí and Bocas del Toro and the east in Coclé and Nevera, ca. 5.5 km N of Hato Chamí (8.5018°N, 81.7689°W),
Colón (Young et al. 1999). 1560 m elev. 9 August 2008. Andreas Hertz and Sebastian Lotz-
Gymnophthalmidae kat. SMF 89519–89521. Ridgetop NE of La Nevera, ca. 5.5 km N
of Hato Chami (8.4954°N, 81.7673°W), 1810 m elev. 18 August
Leposoma southi (Northern Spectacled Lizard). VERAGUAS: 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89522. These
Cerro Mariposa, near Alto de Piedra, ca. 3.5 km W of Santa Fé records, near the type locality of Anolis pseudopachypus, repre-
Herpetological Review 41(4), 2010 521

3. sent the lowest and highest elevational records, respectively, for Viperidae
the species, which was previously known only from 1600 m el-
evation (Köhler et al. 2007). Bothriechis nigroviridis (Black-speckled Palm Viper). COMAR-
CA NGÖBE-BUGLÈ: La Nevera, ca. 5.5 km N of Hato Chamí
Anolis salvini (Salvin’s Anole). CHIRIQUÍ: Jurutungo, near bor- (8.4996°N, 81.77°W), 1650 m elev. 11 May 2008. Andreas Hertz,
der with Costa Rica (8.912°N, 82.7182°W), 1980 m elev. 8 July Leonhard Stadler, Nadim Hamad, and Sebastian Lotzkat. SMF
2008. Andreas Hertz and Sebastian Lotzkat. SMF 89527. This 89534. First record for Comarca Ngöbe-Buglé, and an 80 km
record extends the known elevational distribution of the species E range extension from the formerly easternmost record in Bo-
upward ca. 150 m (Köhler 2008; Savage 2002). quete, Chiriquí (Dunn 1947).
Anolis vittigerus (Garland Anole). VERAGUAS: Río Chilagres, Bothriechis schlegelii (Eyelash Palm Viper). VERAGUAS: Cer-
ca. 9 km NE of Santa Fé (8.5872°N, 81.0363°W), 480 m elev. 3 ro Mariposa, near Alto de Piedra, approx. 3.5 km W of Santa
June 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89528. Fé (8.5157°N, 81.1197°W), 860 m elev. 13 May 2008. Gunther
First record for Veraguas, extending the range ca. 50 km W of the Köhler. SMF 89535. First record for Veraguas, which fills the gap
formerly westernmost record from north of El Copé, Coclé (Poe between Isla Escudo de Veraguas, Comarca Ngöbe-Buglé (ca. 85
and Ibañez 2007). km NW), and El Valle de Anton, Coclé (ca. 105 km E) (Pérez-
Squamata — Snakes Santos 1999).
Colubridae Acknowledgments.—Collecting and exportation permits SE/A-30-08
and SEX/A-108-08 were provided by A. Salazar, Y. Hidalgo, and J. Gar-
Dipsas articulata (American Snail-eater). VERAGUAS: Cerro cía, Autoridad Nacional del Ambiente (ANAM), Panamá City, Panamá.
Negro, ca. 6 km NNW of Santa Fé (8.5756°N, 81.0976°W), 1000 Q. D. Fuenmayora and V. Martínez provided valuable assistance with ac-
m elev. 29 July 2008. Arcadio Carrizo. SMF 89552. Since locali- quisition of permits. An additional permit for the Comarca Ngöbe-Buglé
ty data for the holotype of D. articulata is imprecise (see remarks was issued by A. Montezuma (ANAM), San Félix, Panamá. For field
in Peters 1960), this is the first validated record for Veraguas. The assistance and logistical support, we thank C. O’Shea and S. Abrego,
A. Bennett, P. Yangüez, O. Cáceres and family, M. Piepenbring, and
record reconfirms the species’ occurrence at 1000 m, reported as
the Peña Solís family. A. Carrizo received financial support from the
its upper elevational limit by Solórzano (2004). DAAD. Field work funding to A. Hertz came from FAZIT-Stiftung and
Dipsas temporalis (Temporal Snail-eater). VERAGUAS: Cer- from Studienstiftung des deutschen Volkes to S. Lotzkat.
ro Mariposa, near Alto de Piedra, ca. 3.5 km W of Santa Fé
(8.5107°N, 81.1207°W), 990 m elev. 10 July 2008. Leonhard literature cited
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of Cerro La Campana, the formerly westernmost record in Pan- tion of two species and notes on defensive behaviors in the Dipsadini
amá (Pérez-Santos 1999). Another specimen (SMF 89553) found (Colubridae). Am. Mus. Novit. [3409]:1–47.
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gap between localities in the provinces of Chiriquí and Bocas del lis pentaprion, with the resurrection of A. beckeri and the description
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Urotheca guentheri (Striped Glasstail). VERAGUAS: Cer- –––––, M. Ponce, J. SunYer, and a. BatiSta. 2007. Four new species
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(8.5002°N, 81.1175°W), 1250 m elev. 5 June 2008. Leonhard Panama (Squamata: Polychrotidae). Herpetologica 63:375–391.
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64:92–108.
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Poe, S., and r. iBañez. 2007. A new species of Anolis lizard from the
Cordillera de Talamanca of western Panama. J. Herpetol. 41:263–270. Only a handful of re-
–––––, i. M. latella, M. J. rYan, and e. W. ScHaad. 2009. A new spe- searchers work on South
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works on this group are
Savage, J. M. 2002. The Amphibians and Reptiles of Costa Rica; A Her-
petofauna Between Two Continents, Between Two Seas. Univ. Chi- scarce, and a new book
cago Press, Chicago, Illinois. on this subject is hence
–––––, k. r. liPS, and r. iBañez. 2008. A new species of Celestus from very welcome. It follows
west-central Panama, with consideration of the status of the genera a first volume in the Terra-
of the Anguidae: Diploglossinae (Squamata). Rev. Biol. Tropical log series on South Asian
56:845–859. agamids, that included
–––––, and J. i. Watling. 2008. Not so rare snakes: a revision of the only part of the Draconi-
Geophis sieboldi group (Colubridae: Dipsadinae) in lower Central nae (Manthey 2008), and
America and Colombia. Zool. J. Linn. Soc. 153:561–599.
it aims to list and illustrate
Solórzano, a. 2004. Snakes of Costa Rica: Distribution, Taxonomy, and
Natural History. Instituto Nac. de Biodiversidad (INBio), San José, the remaining taxa that
Costa Rica. were not dealt with in that
uetz, P. 2008. The TIGR Reptile Database. CD ROM edition, August first volume. Taxa newly
2008. http://www.reptile-database.org/ dealt with are all currently
Young, B. e., g. SedagHatkiSH, e. roca, and Q. d. fuenMaYor. 1999. recognized Draconinae
El estatus de la conservación de la herpetofauna de Panamá. Resumen belonging to genera start-
del primer taller internacional sobre la herpetofauna de Panamá. The ing with the letters G to S (Gonocephalus, Harpesaurus, Hyp-
Nature Conservancy y Asociación para la Conservación de la Natura- sicalotes, Japalura, Lophocalotes, Lyriocephalus, Mantheyus,
leza, Panamá City, Panamá.
Mictopholis, Oriocalotes, Otocryptis, Phoxophrys, Psammophi-
lus, Pseudocalotes, Pseudocophotis, Ptyctolaemus, Salea, and Si-
tana), the Leiolepidinae (Leiolepis) and the genus Physignathus
(whose subfamilial position is still unclear), as well as several
undescribed taxa. The book, well bound and with high-quality
glossy paper, is bilingual (English-German). It is divided mainly
into a table of contents (pp. 3–5), an introduction in which the
author gives his point of view on the taxonomic status of various
taxa (pp. 6–8), a section on how to use the book, which provides
brief advice on captive maintenance (pp. 9–11), literature refer-
ences (pp. 12–14), and the main section, the species illustrations
(pp. 16–168). The front and back covers are finely illustrated and
the inner covers provide geographical and political maps of the
area covered.
Herpetological Review 41(4), 2010 523