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Lecture 4. Digestion and Nutrition

                      Major Nutritional Modes
Mode of Nutrition      Energy   Carbon Source               Examples
                       Source
                                Autotrophs
Photoautotrophs     Light       CO2             Photosynthetic prokaryotes,
                                                including cyanobacteria; plants;
                                                certain protists
Chemoautotrophs     Inorganic   CO2             Certain prokaryotes (i.e.
                    chemicals                   Sulfolobus)
                                Heterotrophs
Photoheterotroph    Light       Organic         Certain prokaryotes
                                compounds
Chemoheterotrophs Organic       Organic         Many prokaryotes and protists;
                  compounds     compounds       fungi; animals; parasitic plants
Dietary categories

1. Herbivores
Dietary categories

2. Carnivores
Dietary categories

3. Omnivores
Dietary categories
4. Saprophagous




                                       deer fly (Chrysops callidus)

Woodlice (Trachelipus ratzeburgii)




hermit beetle (Osmoderma eremita)    red ant (Formica pallidefulva )
Feeding Adaptations
Suspension feeder – sifts through food particles in the water




                                 Baleen whale
  Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
•
    bivalve molluscs use
    their gills as feeding
    devices
•
    herring and other suspension-feeding fishes use gill rakers
    to strain plankton
krill


•
    whalebone whales filter out plankton, mainly large
    crustaceans called krill, with whalebone or baleen
Feeding Adaptations
Deposit feeder – eats its way through dirt or sediments
  and extract partially decayed organic material (detritus)
  consumed along with the soil or sediments




 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings   Earthworm
Feeding Adaptations
    Substrate feeder – lives in or on its food source,
      eating its way through the food




                                         Leaf miners
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Feeding Adaptations
    Fluid feeder – sucks nutrient-rich fluids from a living
      host and is considered a parasite




                                                                            Mosquito
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Feeding Adaptations
    Bulk feeder – eats relatively large pieces of food




                                                     Python
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Cephalopod
molluscs have
beak-like jaws
which serve as
tearing devices
•
    chewing mouthparts
    adapted for seizing
    and crushing food

•
    mandibles are strong,
    toothed plates whose
    edges can bite or tear

•
    maxillae hold the
    food and pass it
    toward the mouth
•
    birds lack teeth

•
    bills are often
    provided with
    serrated edges

•
    In some, the upper
    bill is hooked for
    seizing and tearing
    prey
Four types of teeth found in mammals:
   •
     incisors, for biting, cutting, and stripping
    •
        canines, for seizing, piercing, and tearing
    •
        premolars, for grinding and crushing
    •
        molars, for grinding and crushing
•
    an elephant’s tusk is a
    modified upper incisor
•
    used for defense,
    attack, and rooting

•
    a male wild boar has
    modified canines that
    are used as weapons
Types of Digestive System
•
    Incomplete – there is only one opening; no anus




                Paramecium                Hydra
Types of Digestive System
•
    Complete – there is a mouth opening and an anus
Four Main Stages of Food Processing

•
    Ingestion – the act of eating
•
    Digestion – breaking food down into molecules
                small enough for the body to absorb
•
    Absorption – small molecules are taken in
                 by the animal’s cells
•
    Elimination – undigested material passes out of
                  the digestive compartment
•
    digestion is entirely intracellular in
    protozoa and sponges
•
    radiates, turbellarian flatworms,
    and ribbon worms (nemerteans)
    practice both intracellular and
    extracellular digestion
•
    in extracellular digestion, certain
    cells lining the lumen of alimentary
    canals form digestive secretions;
    other cells function in absorption
•
    for arthropods and vertebrates
    digestion is almost entirely
    extracellular
Gut movement
•
    alternate constriction of
    rings of smooth muscle of
    the intestine
•
    constantly divide and
    squeeze contents back
    and forth
•
    for mixing of food

•
    waves of contraction of
    circular muscle behind
    the gut and relaxation in
    front of bolus
•
    sweeps food down the
    gut
Human Digestive
   System Salivary glands
Human Esophagus




   Food is called bolus if it passes
   •


   through the esophagus
Human Stomach
Food is called
•


chyme after it
passes through the
stomach
•
    trypsin, chymotrypsin, and carboxypeptidase
    are secreted in inactive form by the pancreas

    –
        the intestinal enzyme,
        enteropeptidase, converts
        inactive trypsinogen into
        active trypsin

    –
        active trypsin then
        activates the other two
Human Small Intestine
 •
     Made up of three regions:
      1. duodenum
          - Nearest to the stomach; 26 cm in length
          - With Bruner’s glands
              -
                produce mucus-rich alkaline secretion with
                bicarbonates to
                  -
                    protect the duodenum from the acidic content
                    of chyme;
                  -
                    provide alkaline condition for the intestinal
                    enzymes to be active; and
                  -
                    lubricate the intestinal walls
Human Small Intestine
 •
     Made up of three regions:
      2. jejunum
          - Next to duodenum; 2.5 m in length

      3. ileum
          - Last segment of the small intestine; 3.5 m in length
          - Contains Peyer’s patches
               -
                 Organized lymphoid tissues
               -
                 Protects the intestinal lumen from pathogenic
                 bacteria
•
    Food is called chyle once it is in the small intestine
Human Large Intestine
•
    the colon recovers water that has entered the alimentary canal
    as the solvent to various digestive juices

•
    the large intestine harbors a rich flora of mostly harmless
    bacteria

•
    the terminal portion of the colon is called the rectum, where
    feces are stored until they can be eliminated
Regulation of Digestion




CCK - cholecystokinin
•
    the length of the vertebrate digestive system is
    also correlated with diet
•
    ruminants (deer, cattle, and sheep) have the
    most elaborate adaptations for a herbivorous
    diet
Nutritional Requirements

•
    A nutritionally adequate diet satisfies three
    needs:
    –
        fuel (chemical energy) for all the cellular work of the
        body
    –
        organic raw materials animals use in biosynthesis
        (carbon skeletons to make many of their own
        molecules)
    –
        essential nutrients
–
    If the diet of a person or other animal is chronically
    deficient in calories, undernourishment results
–
    an animal whose diet is missing one or more
    essential nutrients is said to be malnourished
–
    marasmus, general undernourishment from a diet
    low in both calories and protein
–
    kwashiorkor, protein malnourishment from a diet
    adequate in calories but deficient in protein
–
    overnourishment or obesity results from excessive
    food intake
•
    in mammals, a hormone called leptin,
    produced by adipose cells, is a key player in a
    complex feedback mechanism regulating fat
    storage and use
Appetite-regulating hormones
•
    ghrelin (stomach wall)
    –
        triggers feelings of hunger as mealtimes approach
•
    leptin (adipose tissue)
    –
        suppresses appetite
•
    PYY or peptide-tyrosine-tyrosine (small intestine)
    –
        after meals
    –
        appetite suppressant; counters ghrelin
•
    insulin (pancreas)
    –
        a rise in blood sugar level after a meal
•
    animals require 20 amino acids to make proteins
•
    essential amino acids must be obtained from
    food in prefabricated form

    –
        eight amino acids are essential in the adult human
        (phenylalanine, lysine, isoleucine, leucine, valine,
        methionine, tryptophan, and threonine) with
        histidine and arginine essential for normal growth of
        children
•
    protein deficiency from a vegetarian diet can
    be avoided by eating a combination of plant
    foods that complement each other to supply all
    essential amino acids
•
    vitamins are organic molecules required in the
    diet in small quantities
•
    13 vitamins essential to humans have been
    identified
•
    water-soluble vitamins generally function as
    co-enzymes
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
•
    Minerals are simple inorganic nutrients, usually
    required in small amounts
    –
        humans and other vertebrates require large
        quantities of calcium and phosphorus for the
        construction and maintenance of bone
    –
        iron is a component of the cytochromes and of
        hemoglobin
    –
        while sodium, potassium, and chloride have a
        major influence on the osmotic balance between
        cells and the interstitial fluids, excess consumption
        of salt (sodium chloride) is harmful
Lecture 4 digestion and nutrition 2nd sem 2008-2009

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Lecture 4 digestion and nutrition 2nd sem 2008-2009

  • 1. Lecture 4. Digestion and Nutrition Major Nutritional Modes Mode of Nutrition Energy Carbon Source Examples Source Autotrophs Photoautotrophs Light CO2 Photosynthetic prokaryotes, including cyanobacteria; plants; certain protists Chemoautotrophs Inorganic CO2 Certain prokaryotes (i.e. chemicals Sulfolobus) Heterotrophs Photoheterotroph Light Organic Certain prokaryotes compounds Chemoheterotrophs Organic Organic Many prokaryotes and protists; compounds compounds fungi; animals; parasitic plants
  • 5. Dietary categories 4. Saprophagous deer fly (Chrysops callidus) Woodlice (Trachelipus ratzeburgii) hermit beetle (Osmoderma eremita) red ant (Formica pallidefulva )
  • 6. Feeding Adaptations Suspension feeder – sifts through food particles in the water Baleen whale Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 7. bivalve molluscs use their gills as feeding devices
  • 8. herring and other suspension-feeding fishes use gill rakers to strain plankton
  • 9. krill • whalebone whales filter out plankton, mainly large crustaceans called krill, with whalebone or baleen
  • 10. Feeding Adaptations Deposit feeder – eats its way through dirt or sediments and extract partially decayed organic material (detritus) consumed along with the soil or sediments Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Earthworm
  • 11. Feeding Adaptations Substrate feeder – lives in or on its food source, eating its way through the food Leaf miners Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 12. Feeding Adaptations Fluid feeder – sucks nutrient-rich fluids from a living host and is considered a parasite Mosquito Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 13. Feeding Adaptations Bulk feeder – eats relatively large pieces of food Python Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 15. chewing mouthparts adapted for seizing and crushing food • mandibles are strong, toothed plates whose edges can bite or tear • maxillae hold the food and pass it toward the mouth
  • 16. birds lack teeth • bills are often provided with serrated edges • In some, the upper bill is hooked for seizing and tearing prey
  • 17. Four types of teeth found in mammals: • incisors, for biting, cutting, and stripping • canines, for seizing, piercing, and tearing • premolars, for grinding and crushing • molars, for grinding and crushing
  • 18. an elephant’s tusk is a modified upper incisor • used for defense, attack, and rooting • a male wild boar has modified canines that are used as weapons
  • 19.
  • 20. Types of Digestive System • Incomplete – there is only one opening; no anus Paramecium Hydra
  • 21. Types of Digestive System • Complete – there is a mouth opening and an anus
  • 22. Four Main Stages of Food Processing • Ingestion – the act of eating • Digestion – breaking food down into molecules small enough for the body to absorb • Absorption – small molecules are taken in by the animal’s cells • Elimination – undigested material passes out of the digestive compartment
  • 23. digestion is entirely intracellular in protozoa and sponges
  • 24. radiates, turbellarian flatworms, and ribbon worms (nemerteans) practice both intracellular and extracellular digestion • in extracellular digestion, certain cells lining the lumen of alimentary canals form digestive secretions; other cells function in absorption • for arthropods and vertebrates digestion is almost entirely extracellular
  • 25. Gut movement • alternate constriction of rings of smooth muscle of the intestine • constantly divide and squeeze contents back and forth • for mixing of food • waves of contraction of circular muscle behind the gut and relaxation in front of bolus • sweeps food down the gut
  • 26.
  • 27. Human Digestive System Salivary glands
  • 28. Human Esophagus Food is called bolus if it passes • through the esophagus
  • 30. Food is called • chyme after it passes through the stomach
  • 31. trypsin, chymotrypsin, and carboxypeptidase are secreted in inactive form by the pancreas – the intestinal enzyme, enteropeptidase, converts inactive trypsinogen into active trypsin – active trypsin then activates the other two
  • 32. Human Small Intestine • Made up of three regions: 1. duodenum - Nearest to the stomach; 26 cm in length - With Bruner’s glands - produce mucus-rich alkaline secretion with bicarbonates to - protect the duodenum from the acidic content of chyme; - provide alkaline condition for the intestinal enzymes to be active; and - lubricate the intestinal walls
  • 33. Human Small Intestine • Made up of three regions: 2. jejunum - Next to duodenum; 2.5 m in length 3. ileum - Last segment of the small intestine; 3.5 m in length - Contains Peyer’s patches - Organized lymphoid tissues - Protects the intestinal lumen from pathogenic bacteria
  • 34.
  • 35. Food is called chyle once it is in the small intestine
  • 36. Human Large Intestine • the colon recovers water that has entered the alimentary canal as the solvent to various digestive juices • the large intestine harbors a rich flora of mostly harmless bacteria • the terminal portion of the colon is called the rectum, where feces are stored until they can be eliminated
  • 37.
  • 38.
  • 39.
  • 40. Regulation of Digestion CCK - cholecystokinin
  • 41. the length of the vertebrate digestive system is also correlated with diet
  • 42. ruminants (deer, cattle, and sheep) have the most elaborate adaptations for a herbivorous diet
  • 43. Nutritional Requirements • A nutritionally adequate diet satisfies three needs: – fuel (chemical energy) for all the cellular work of the body – organic raw materials animals use in biosynthesis (carbon skeletons to make many of their own molecules) – essential nutrients
  • 44. If the diet of a person or other animal is chronically deficient in calories, undernourishment results – an animal whose diet is missing one or more essential nutrients is said to be malnourished – marasmus, general undernourishment from a diet low in both calories and protein – kwashiorkor, protein malnourishment from a diet adequate in calories but deficient in protein – overnourishment or obesity results from excessive food intake
  • 45. in mammals, a hormone called leptin, produced by adipose cells, is a key player in a complex feedback mechanism regulating fat storage and use
  • 46. Appetite-regulating hormones • ghrelin (stomach wall) – triggers feelings of hunger as mealtimes approach • leptin (adipose tissue) – suppresses appetite • PYY or peptide-tyrosine-tyrosine (small intestine) – after meals – appetite suppressant; counters ghrelin • insulin (pancreas) – a rise in blood sugar level after a meal
  • 47. animals require 20 amino acids to make proteins • essential amino acids must be obtained from food in prefabricated form – eight amino acids are essential in the adult human (phenylalanine, lysine, isoleucine, leucine, valine, methionine, tryptophan, and threonine) with histidine and arginine essential for normal growth of children
  • 48. protein deficiency from a vegetarian diet can be avoided by eating a combination of plant foods that complement each other to supply all essential amino acids
  • 49. vitamins are organic molecules required in the diet in small quantities • 13 vitamins essential to humans have been identified • water-soluble vitamins generally function as co-enzymes
  • 50. Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 51. Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
  • 52. Minerals are simple inorganic nutrients, usually required in small amounts – humans and other vertebrates require large quantities of calcium and phosphorus for the construction and maintenance of bone – iron is a component of the cytochromes and of hemoglobin – while sodium, potassium, and chloride have a major influence on the osmotic balance between cells and the interstitial fluids, excess consumption of salt (sodium chloride) is harmful