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The effect of grazing intensity on grasslands and cattle performance in south-central North Dakota
                                                                                                                                                                            Bob Patton, Paul Nyren, and Anne Nyren
                                                                                                                                                    North Dakota State University - Central Grasslands Research Extension Center, Streeter ND
                    Abstract                                                                                                                   Forage Production                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    Livestock Response                                                                                                                                                                                          Economics
    A long-term grazing intensity study began                  On the loamy ecological site, the greatest                                                                                                                                                          Some species that increase under grazing:                                                                                                                                                                                                                Some species favored by moderate grazing:                                                                                                                                                Average daily gain and condition scores                                                                                                                                              If cattle prices were constant, then return/ton
at CGREC in 1989 to determine the                              forage production is on the light treatment.                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          decrease as grazing intensity increases. Gain                                                                                                                                        would peak at a stocking rate somewhere
                                                                                                                                                                                                                                                                   Pascopyrum smithii - western wheatgrass                                                                                                                                                                                                                  Oligoneuron rigidum - stiff goldenrod                                                                                                                                                                                                                                                                                                                                         below maximum gain/ton, with the exact point
ecological and economic effects of season-                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                           per ton of forage initially goes up as grazing
                                                                                         Average above ground biomass production by grazing treatment on loamy ecological sites from 1992 to 2010.
                                                                                                                                                                                                                                                                   Carex inops ssp. heliophila - sun sedge                                                                                                                                                                                                                  Ambrosia psilostachya - western ragweed                                                                                                                                                                                                                                                                                                                                       depending on carrying costs. The change in
long cattle grazing at different intensities. Five                                                                                                                                          Above ground biomass (lbs/acre)                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          intensity increases, but there is a point
                                                                                                                                                              Beginning                          Middle of                Peak                     End of          Nassella viridula - green needlegrass                                                                                                                                                                                                                    Solidago canadensis - Canada goldenrod                                                                                                                                                                                                                                                                                                                                        cattle prices over the season determines the
treatments - no grazing, light, moderate,                                                Treatment                                                            of season                           season                  yield                    season                                                                                                                                                                                                                                                                                                                                                                                                                                                            beyond which gain per ton decreases with
                                                                                                                                                                                                                                                                   Achillea millefolium - western yarrow                                                                                                                                                                                                                    Glycyrrhiza lepidota - wild licorice                                                                                                                                                                                                                                                                                                                                          stocking rate with the maximum return/ton.
heavy, and extreme grazing - are each                                                    Ungrazed                                                              1,253 b1                          2,448 b                 2,728 c                   2,592 c                                                                                                                                                                                                                                                                                                                                                                                                                                                           increasing grazing intensity.
                                                                                         Light                                                                 1,314 a                           2,765 a                 3,168 a                  3,071 a
                                                                                                                                                                                                                                                                   Bouteloua gracilis - blue grama                                                                                                                                                                                                                          Solidago mollis - soft goldenrod                                                                                                                                                                                                                                                                                                                                              The stocking rate with the maximum
replicated three times. Pastures are                                                                                                                                                                                                                               Taraxacum officinale - common dandelion                                                                                                                                                                                                                  Carex lanuginosa - wooly sedge                                                                                                                                                                       Average daily gains, gains per acre, gains per ton of forage and condition scores from different
approximately 30 acres each. The no grazing
                                                                                         Moderate                                                              1,178 c                           2,535 b                 2,941 b                  2,844 b
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 stocking intensities.                                                                                                                                                    return/ton over the last 20 years would be
                                                                                         Heavy                                                                     901 d                          2,148 c                2,421 d                  2,339 d          Artemisia frigida - fringed sagewort                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   1.74 AUM/ton, with an average annual return
treatment consists of six 0.3-acre exclosures.                                           Extreme                                                                   732 e                         1,821 d                 2,189 e                  2,150 d
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                            An example: Oligoneuron rigidum
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   Average Daily Gains (lbs./head/day)

Pastures are stocked so that when the cattle                                             LSD (0.05)                                                                59                                 142                 183                         196
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Desired Grazing                                                                                                                    Average 1991-
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          of $28.24/ton.
                                                                                         1
                                                                                                                                                                                                                                                                   An example: Potentilla pensylvanica                                                                                                                                                                                                                                                                                                                                                                                                                                 Intensity                                   2006               2007            2008                       2009          2010       2010

are removed in the fall, about 65, 50, 35, and                                                               Means in the same column followed by the same letter are not significantly different at P=0.05.
                                                                                                                                                                                                                                                                                                                                                               30                                                                                                                                                                                                       500
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Light                                        0.57              1.36            1.75a1                     2.05a         1.54        1.39a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Moderate                                     0.62              1.22            1.58ab                     1.99a         1.29        1.28a
20% of the forage produced in an average                    On the loamy overflow ecological site,                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     Heavy                                        0.48              1.33            1.35b                      1.48b         1.09        1.10b
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                  140
year remains on the light, moderate, heavy,                 forage production does not differ between the                                                                                                                                                                                                                                                                                                                                                                                               E                                                                                                                          M
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Extreme                                      0.13              1.16            0.95c                      1.09b         1.02        0.80c
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                  120                                                                                                                                        2007
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       LSD (0.05)                                   NS2               NS              0.38                       0.42          NS          0.14




                                                                                                                                                                                                                                                                                                        Percent frequency of occurrence in 25 X 25 cm frames
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             M
and extreme grazing treatments respectively.                light, moderate, and heavy treatments, but
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        400
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          Average Gain (lbs./acre)                                                                100
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               Average 1991-
                                                            ungrazed and extreme produce significantly                                                                                                                                                                                                                                                         20
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    2006              2007            2008                       2009          2010            2010
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        80
   The two most common ecological sites,                    less forage.                                                                                                                                                                                                                                                                                                                                                                                                                       E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        300
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Light                                        11.01             44.41c          39.73b                     47.37b         47.58      30.02c
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        60
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               up
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   pe
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     rc
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             .i.
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          20                       of
loamy and loamy overflow, are monitored.                                                                                                                                                                                                                                                                                                                                                                                                                                                            H                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   2008                06                          20




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    Plants/M2
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Moderate                                     20.82             69.27bc         68.61ab                    90.63a         68.95      55.19b                                       40




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          Dollars/Ton
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             -yr
                                                                                                 Average above ground biomass production by grazing treatment on loamy overflow ecological sites from 1993 to 2010.                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   20-yr. a                                                  .a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               ve.                                                  ve
Forage production on the loamy site is highest                                                                                                                                               Above ground biomass (lbs/acre)                                                                                                                                                                       E                      E
                                                                                                                                                                                                                                                                                                                                                                                                                          H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                   E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 E                                                                                                                                                                                                                                                     Heavy
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       Extreme
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    20.77
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     7.48
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                      107.47ab
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                      122.96a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                      82.15a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                      76.10a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 92.72a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 90.79a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                84.55
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               104.70
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                           76.45a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                           81.42a
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        20                                                                                                                               .
                                                                                                                                                                                                                                                                                                                                                                                                         H
                                                                                                                                                                                                                                                                                                                                                                                                         E                                                                               E
under the light grazing treatment. On the                                                        Treatment
                                                                                                                                                               Beginning
                                                                                                                                                               of season
                                                                                                                                                                                                 Middle of
                                                                                                                                                                                                  season
                                                                                                                                                                                                                          Peak
                                                                                                                                                                                                                          yield
                                                                                                                                                                                                                                                   End of
                                                                                                                                                                                                                                                   season
                                                                                                                                                                                                                                                                                                                                                               10
                                                                                                                                                                                                                                                                                                                                                                                                   H
                                                                                                                                                                                                                                                                                                                                                                                                              H
                                                                                                                                                                                                                                                                                                                                                                                                                    H
                                                                                                                                                                                                                                                                                                                                                                                                                    E                              E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        200
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             H                         LSD (0.05)                                   NS                42.67           29.04                      34.31         NS          10.30                                         0
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                          200
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                               9
loamy overflow site, production does not differ                                                  Ungrazed                                                      1,002 b      1
                                                                                                                                                                                                  3,205 c                3,350 b                   2,928 b
                                                                                                                                                                                                                                                                                                                                                                                                              E                    E
                                                                                                                                                                                                                                                                                                                                                                                                                                           E
                                                                                                                                                                                                                                                                                                                                                                                                                                                           E                                                                                                                                                                       U
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     M                 E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             E                                                                                                       Average Gain (lbs./ton of forage)
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                           Average 1991-
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    -20
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 low
                                                                                                                                                                                                                                                                                                                                                                                                         M                                                             E                                     H                                                                                                                                                                                               L                                                                                                                                                                                                                                                                                                          er
between light, moderate, and heavy, but                                                          Light                                                          1,156 a                           3,890 a                4,173 a                   3,936 a                                                                                                                                                                     H
                                                                                                                                                                                                                                                                                                                                                                                                                                       H
                                                                                                                                                                                                                                                                                                                                                                                                                                               H
                                                                                                                                                                                                                                                                                                                                                                                                                                               E   H
                                                                                                                                                                                                                                                                                                                                                                                                                                                       H               H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                       M      H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    M
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        100
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   L
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        M
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             M
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 H
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     E
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                       H     M                                                                      2006              2007            2008                       2009          2010        2010
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    -40                                                                                                      c.i
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   .o
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        f2
                                                                                                 Moderate                                                       1,230 a                           3,666 ab               4,116 a                   3,994 a                                                                                                                                                                                                                                                                                                                                                                                                                   E   E          E    E                                                                                                                                                                                                                  -60
                                                                                                                                                                                                                                                                                                                                                                                              E               M     M     M                                                                                                                                                                                                                              M                                  H                                          Light                                        9.83              23.19c          27.11c                     33.80b        19.01c      19.53d
ungrazed and extreme treatments produce                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   0-




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                             Moderate




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                            Optimum
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 H




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                 20
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     E                                 L




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                Extreme
                                                                                                                                                                                                                                                                                                                                                                                                                                                       M                                                                                                                                                                     L                               L   E   H   E   M   M                     M                                                                                                                                                                                                                                                                                                                                      yr.
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     L




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                              Heavy
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                        H                        H       M




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                    10
                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                     M




                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                                   Light

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The effect of grazing intensity on the grasslands and cattle performance v8

  • 1. The effect of grazing intensity on grasslands and cattle performance in south-central North Dakota Bob Patton, Paul Nyren, and Anne Nyren North Dakota State University - Central Grasslands Research Extension Center, Streeter ND Abstract Forage Production Livestock Response Economics A long-term grazing intensity study began On the loamy ecological site, the greatest Some species that increase under grazing: Some species favored by moderate grazing: Average daily gain and condition scores If cattle prices were constant, then return/ton at CGREC in 1989 to determine the forage production is on the light treatment. decrease as grazing intensity increases. Gain would peak at a stocking rate somewhere Pascopyrum smithii - western wheatgrass Oligoneuron rigidum - stiff goldenrod below maximum gain/ton, with the exact point ecological and economic effects of season- per ton of forage initially goes up as grazing Average above ground biomass production by grazing treatment on loamy ecological sites from 1992 to 2010. Carex inops ssp. heliophila - sun sedge Ambrosia psilostachya - western ragweed depending on carrying costs. The change in long cattle grazing at different intensities. Five Above ground biomass (lbs/acre) intensity increases, but there is a point Beginning Middle of Peak End of Nassella viridula - green needlegrass Solidago canadensis - Canada goldenrod cattle prices over the season determines the treatments - no grazing, light, moderate, Treatment of season season yield season beyond which gain per ton decreases with Achillea millefolium - western yarrow Glycyrrhiza lepidota - wild licorice stocking rate with the maximum return/ton. heavy, and extreme grazing - are each Ungrazed 1,253 b1 2,448 b 2,728 c 2,592 c increasing grazing intensity. Light 1,314 a 2,765 a 3,168 a 3,071 a Bouteloua gracilis - blue grama Solidago mollis - soft goldenrod The stocking rate with the maximum replicated three times. Pastures are Taraxacum officinale - common dandelion Carex lanuginosa - wooly sedge Average daily gains, gains per acre, gains per ton of forage and condition scores from different approximately 30 acres each. The no grazing Moderate 1,178 c 2,535 b 2,941 b 2,844 b stocking intensities. return/ton over the last 20 years would be Heavy 901 d 2,148 c 2,421 d 2,339 d Artemisia frigida - fringed sagewort 1.74 AUM/ton, with an average annual return treatment consists of six 0.3-acre exclosures. Extreme 732 e 1,821 d 2,189 e 2,150 d An example: Oligoneuron rigidum Average Daily Gains (lbs./head/day) Pastures are stocked so that when the cattle LSD (0.05) 59 142 183 196 Desired Grazing Average 1991- of $28.24/ton. 1 An example: Potentilla pensylvanica Intensity 2006 2007 2008 2009 2010 2010 are removed in the fall, about 65, 50, 35, and Means in the same column followed by the same letter are not significantly different at P=0.05. 30 500 Light 0.57 1.36 1.75a1 2.05a 1.54 1.39a Moderate 0.62 1.22 1.58ab 1.99a 1.29 1.28a 20% of the forage produced in an average On the loamy overflow ecological site, Heavy 0.48 1.33 1.35b 1.48b 1.09 1.10b 140 year remains on the light, moderate, heavy, forage production does not differ between the E M Extreme 0.13 1.16 0.95c 1.09b 1.02 0.80c 120 2007 LSD (0.05) NS2 NS 0.38 0.42 NS 0.14 Percent frequency of occurrence in 25 X 25 cm frames M and extreme grazing treatments respectively. light, moderate, and heavy treatments, but 400 Average Gain (lbs./acre) 100 Average 1991- ungrazed and extreme produce significantly 20 H 2006 2007 2008 2009 2010 2010 80 The two most common ecological sites, less forage. E E 300 Light 11.01 44.41c 39.73b 47.37b 47.58 30.02c 60 up pe rc .i. 20 of loamy and loamy overflow, are monitored. H 2008 06 20 Plants/M2 Moderate 20.82 69.27bc 68.61ab 90.63a 68.95 55.19b 40 Dollars/Ton -yr Average above ground biomass production by grazing treatment on loamy overflow ecological sites from 1993 to 2010. 20-yr. a .a ve. ve Forage production on the loamy site is highest Above ground biomass (lbs/acre) E E H E E Heavy Extreme 20.77 7.48 107.47ab 122.96a 82.15a 76.10a 92.72a 90.79a 84.55 104.70 76.45a 81.42a 20 . H E E under the light grazing treatment. On the Treatment Beginning of season Middle of season Peak yield End of season 10 H H H E E H 200 H LSD (0.05) NS 42.67 29.04 34.31 NS 10.30 0 200 9 loamy overflow site, production does not differ Ungrazed 1,002 b 1 3,205 c 3,350 b 2,928 b E E E E U H M E E Average Gain (lbs./ton of forage) Average 1991- -20 low M E H L er between light, moderate, and heavy, but Light 1,156 a 3,890 a 4,173 a 3,936 a H H H E H H H M H M 100 H L M M H H E H M 2006 2007 2008 2009 2010 2010 -40 c.i .o f2 Moderate 1,230 a 3,666 ab 4,116 a 3,994 a E E E E -60 E M M M M H Light 9.83 23.19c 27.11c 33.80b 19.01c 19.53d ungrazed and extreme treatments produce 0- Moderate Optimum H 20 E L Extreme M L L E H E M M M yr. L Heavy H H M 10 M Light