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Sensory Systems
• Vision
• Hearing
• Taste
• Smell
• Equilibrium
• Somatic Senses
Sensory Systems
• Somatic sensory
• General – transmit impulses from skin, skeletal muscles,
and joints
• Special senses - hearing, balance, vision
• Visceral sensory
• Transmit impulses from visceral organs
• Special senses - olfaction (smell), gustation (taste)
• Stimulus - energy source
• Internal
• External
• Receptors
• Sense organs - structures specialized to respond to
stimuli
• Transducers - stimulus energy converted into
action potentials
• Conduction
• Afferent pathway
• Nerve impulses to the CNS
• Translation
• CNS integration and information processing
• Sensation and perception – your reality
Properties of Sensory Systems
Sensory Pathways
• Stimulus as physical energy  sensory receptor acts as a transducer
• Stimulus > threshold  action potential to CNS
• Integration in CNS  cerebral cortex or acted on subconsciously
Classification by Function (Stimuli)
• Mechanoreceptors – respond to touch, pressure, vibration, stretch, and itch
• Thermoreceptors – sensitive to changes in temperature
• Photoreceptors – respond to light energy (e.g., retina)
• Chemoreceptors – respond to chemicals (e.g., smell, taste, changes in
blood chemistry)
• Nociceptors – sensitive to pain-causing stimuli
• Osmoreceptors – detect changes in concentration of solutes, osmotic
activity
• Baroreceptors – detect changes in fluid pressure
Classification by Location
• Exteroceptors – sensitive to stimuli arising from outside the body
• Located at or near body surfaces
• Include receptors for touch, pressure, pain, and temperature
• Interoceptors – (visceroceptors) receive stimuli from internal viscera
• Monitor a variety of stimuli
• Proprioceptors – monitor degree of stretch
• Located in musculoskeletal organs
Classification by Structure
• General somatic – include touch,
pain, vibration, pressure, temperature
• Proprioceptive – detect stretch in
tendons and muscle provide
information on body position,
orientation and movement of body in
space
Somatic Senses
Somatic Receptors
• Divided into two groups
• Free or Unencapsulated nerve endings
• Encapsulated nerve endings - consist of one or more neural
end fibers enclosed in connective tissue
Free Nerve Endings
• Abundant in epithelia and underlying connective tissue
• Nociceptors - respond to pain
• Thermoreceptors - respond to temperature
• Two specialized types of free nerve endings
• Merkel discs – lie in the epidermis, slowly adapting receptors for light touch
• Hair follicle receptors – Rapidly adapting receptors that wrap around hair follicles
Encapsulated Nerve Endings
• Meissner’s corpuscles
• Spiraling nerve ending surrounded by Schwann cells
• Occur in the dermal papillae of hairless areas of the skin
• Rapidly adapting receptors for discriminative touch
• Pacinian corpuscles
• Single nerve ending surrounded by layers of flattened Schwann cells
• Occur in the hypodermis
• Sensitive to deep pressure – rapidly adapting receptors
• Ruffini’s corpuscles
• Located in the dermis and respond to pressure
• Monitor continuous pressure on the skin – adapt slowly
Encapsulated Nerve Endings - Proprioceptors
• Monitor stretch in locomotory organs
• Three types of proprioceptors
• Muscle spindles – monitors the changing length of a muscle, imbedded in the
perimysium between muscle fascicles
• Golgi tendon organs – located near the muscle-tendon junction, monitor tension
within tendons
• Joint kinesthetic receptors - sensory nerve endings within the joint capsules, sense
pressure and position
Muscle Spindle & Golgi Tendon Organ
Special Senses
Figure 10-4: Sensory pathways
• Taste, smell, sight,
hearing, and balance
• Localized – confined to
the head region
• Receptors are not free
endings of sensory
neurons but specialized
receptor cells
Anatomy of the Eyeball
• Function of the eyeball
• Protect and support the photoreceptors
• Gather, focus, and process light into precise images
• External walls – composed of three tunics (layers)
• Internal cavity – contains fluids (humors)
The Fibrous Layer
• Most external layer of the eyeball
• Cornea
• Anterior one-sixth of the fibrous tunic
• Composed of stratified Squamous externally, simple squamous internally
• Refracts (bends) light
• Sclera
• Posterior five-sixths of the tunic
• White, opaque region composed of dense irregular connective tissue Provides
shape and an anchor for eye muscles,
• Scleral venous sinus – allows aqueous humor to drain
The Vascular Layer
• Middle layer consists of choroid, ciliary body, and iris
• Iris and Pupil
• Composed of smooth muscle, melanocytes, and blood vessels that forms the colored portion of
the eye.
• Function: It regulates the amount of light entering the eye through the pupil.
• It is attached to the ciliary body.
• Pupil is the opening in center of iris through which light enters the eye
• Ciliary body
• Composed of a ring of muscle called ciliary muscle and ciliary processes which are folds located
at the posterior surface of ciliary bodies
• Suspensory ligaments attach to these processes
• Function: secretes the aqueous humor
• The suspensory ligaments position the lens so that light passing through the pupil passes through
the center of the lens of the eye.
The Vascular Layer
• Choroid - vascular layer in the wall of the eye.
• Dark brown (pigmented) membrane with melanocytes that lines most of
the internal surface of the sclera. Has lots of blood vessels
• Lines most of the interior of the sclera.
• Extends from the ciliary body to the lens.
• Corresponds to arachnoid and pia mater
• Functions:
• Delivers oxygen and nutrients to the retina.
• Absorb light rays so that the light rays are not reflected within the eye
The Inner Layer (Retina)
• Retina is the innermost layer of the eye lining the posterior cavity
• The retina contains 2 layers:
• Pigmented layer made of a single layer of melanocytes, absorbs light after it
passes through the neural layer
• Neural layer – sheet of nervous tissue, contains three main types of neurons
• Photoreceptor cells
• Bipolar cells
• Ganglion cells
Photoreceptors
• Two main types
• Rod cells
• More sensitive to light
• Allow vision in dim light
• In periphery
• Cone cells
• Operate best in bright light
• High-acuity
• Color vision – blue, green, red cones
• Concentrated in fovea
Regional Specializations of the Retina
• Ora serrata retinae
• Neural layer ends at the posterior margin of the ciliary body
• Pigmented layer covers ciliary body and posterior surface of the iris
• Macula lutea – contains mostly cones
• Fovea centralis – contains only cones
• Region of highest visual acuity
• Optic disc – blind spot
The Lens
• A thick, transparent, biconvex disc
• Held in place by its ciliary zonule
• Lens epithelium – covers anterior surface of the
lens
The Eye as an Optical Device
• Structures in the eye bend light rays
• Light rays converge on the retina at a single focal point
• Light bending structures (refractory media)
• The lens, cornea, and humors
• Accommodation – curvature of the lens is adjustable
• Allows for focusing on nearby objects
Internal Chambers and Fluids
Figure 16.8
Internal Chambers and Fluids
• Anterior segment
• Divided into anterior and posterior chambers
• Anterior chamber – between the cornea and iris
• Posterior chamber – between the iris and lens
• Filled with aqueous humor
• Renewed continuously
• Formed as a blood filtrate
• Supplies nutrients to the lens and cornea
Internal Chambers and Fluids
• The lens and ciliary zonules divide the eye
• Posterior segment (cavity)
• Filled with vitreous humor - clear, jelly-like substance
• Transmits light
• Supports the posterior surface of the lens
• Helps maintain intraocular pressure
Accessory Structures of the Eye
• Eyebrows – coarse hairs on
the superciliary arches
• Eyelids (palpebrae)
• Separated by the palpebral
fissure
• Meet at the medial and
lateral angles (canthi)
• Conjunctiva – transparent
mucous membrane
• Palpebral conjunctiva
• Bulbar (ocular) conjunctiva
• Conjunctival sac
• Moistens the eye
Figure 16.5a
Accessory Structures of the Eye
• Lacrimal apparatus –
keeps the surface of the
eye moist
• Lacrimal gland –
produces lacrimal fluid
• Lacrimal sac – fluid
empties into nasal
cavity
Figure 16.5b
Extrinsic Eye Muscles
Figure 16.6a, b
• Six muscles that control movement of the eye
• Originate in the walls of the orbit
• Insert on outer surface of the eyeball
Visual Pathways to the Cerebral Cortex
• Pathway begins at the retina
• Light activates photoreceptors
• Photoreceptors signal bipolar cells
• Bipolar cells signal ganglion cells
• Axons of ganglion cells exit eye as the optic nerve
• Optic nerve
• Optic chiasm
• Optic tract
• Thalamus
• Visual cortex
• Other pathways
include the
midbrain and
diencephalon
Vision Integration / Pathway
Figure 10-29b, c: Neural pathways for vision and the papillary reflex
The Ear: Hearing and Equilibrium
• The ear – receptor organ for hearing and
equilibrium
• Composed of three main regions
• Outer ear – functions in hearing
• Middle ear – functions in hearing
• Inner ear – functions in both hearing and
equilibrium
The Outer (External) Ear
• Auricle (pinna) - helps direct sounds
• External acoustic meatus
• Lined with skin
• Contains hairs, sebaceous glands, and ceruminous glands
• Tympanic membrane
• Forms the boundary between the external and middle ear
The Middle Ear
• The tympanic cavity
• A small, air-filled space
• Located within the petrous portion of the temporal bone
• Medial wall is penetrated by
• Oval window
• Round window
• Pharyngotympanic tube (auditory or eustachian tube) - Links the middle ear
and pharynx
Figure 16.17
The Middle Ear
• Ear ossicles – smallest
bones in the body
• Malleus – attaches to
the eardrum
• Incus – between the
malleus and stapes
• Stapes – vibrates
against the oval
window
The Inner (Internal) Ear
• Inner ear – also called the
labyrinth
• Bony labyrinth – a cavity
consisting of three parts
• Semicircular canals
• Vestibule
• Cochlea
• Bony labyrinth is filled
with perilymph
The Membranous Labyrinth
Figure 16.18
• Membranous labyrinth - series of membrane-walled sacs and ducts
• Fit within the bony labyrinth
• Consists of three main parts
• Semicircular ducts
• Utricle and saccule
• Cochlear duct
• Filled with a clear fluid – endolymph
The Cochlea
• A spiraling chamber in the bony labyrinth
• Coils around a pillar of bone – the modiolus
• Spiral lamina – a spiral of bone in the modiolus
• The cochlear nerve runs through the core of the modiolus
The Cochlea
• The cochlear duct (scala media) – contains receptors for hearing
• Lies between two chambers
• The scala vestibuli
• The scala tympani
• The vestibular membrane – the roof of the cochlear duct
• The basilar membrane – the floor of the cochlear duct
The Cochlea
• The cochlear duct (scala media) – contains receptors for
hearing
• Organ of Corti – the receptor epithelium for hearing
• Consists of hair cells (receptor cells)
The Role of the Cochlea in Hearing
Figure 16.20
Auditory Pathway from the Organ of Corti
• The ascending
auditory pathway
• Transmits
information from
cochlear receptors
to the cerebral
cortex
Figure 16.23
The Vestibule
• Utricle and saccule – suspended in perilymph
• Two egg-shaped parts of the membranous labyrinth
• House the macula – a spot of sensory epithelium
• Macula – contains receptor cells
• Monitor the position of the head when the head is still
• Contains columnar supporting cells
• Receptor cells – called hair cells
• Synapse with the vestibular nerve
Anatomy and Function of the Maculae
Figure 16.21b
The Semicircular Canals
• Lie posterior and lateral to the vestibule
• Anterior and posterior semicircular canals lie in the vertical plane at
right angles
• Lateral semicircular canal lies in the horizontal plane
The Semicircular Canals
• Semicircular duct – snakes through each semicircular canal
• Membranous ampulla – located within bony ampulla
• Houses a structure called a crista ampullaris
• Cristae contain receptor cells of rotational acceleration
• Epithelium contains supporting cells and receptor hair cells
Structure and Function of the Crista
Ampullaris
Figure 16.22b
The Chemical Senses: Taste and Smell
• Taste – gustation
• Smell – olfaction
• Receptors – classified as chemoreceptors
• Respond to chemicals
Taste – Gustation
• Taste receptors
• Occur in taste buds
• Most are found on the surface of the
tongue
• Located within tongue papillae
• Two types of papillae (with taste buds)
• Fungiform papillae
• Circumvallate papillae
Taste Buds
• Collection of 50 –100 epithelial cells
• Contain three major cell types (similar
in all special senses)
• Supporting cells
• Gustatory cells
• Basal cells
• Contain long microvilli – extend
through a taste pore
Taste Sensation and the Gustatory Pathway
• Four basic qualities of taste
• Sweet, sour, salty, and bitter
• A fifth taste – umami, “deliciousness”
• No structural difference among taste buds
Gustatory Pathway from Taste Buds
Figure 16.2
• Taste information reaches the
cerebral cortex
• Primarily through the facial
(VII) and glossopharyngeal
(IX) nerves
• Some taste information
through the vagus nerve (X)
• Sensory neurons synapse in
the medulla
• Located in the solitary
nucleus
• Olfactory epithelium with olfactory receptors, supporting cells, basal cells
• Olfactory receptors are modified neurons
• Surfaces are coated with secretions from olfactory glands
• Olfactory reception involves detecting dissolved chemicals as they interact with
odorant binding proteins
Smell (Olfaction)
Olfactory Receptors
• Bipolar sensory neurons located within olfactory epithelium
• Dendrite projects into nasal cavity, terminates in cilia
• Axon projects directly up into olfactory bulb of cerebrum
• Olfactory bulb projects to olfactory cortex, hippocampus, and amygdaloid
nuclei

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Sensory organs

  • 1. Sensory Systems • Vision • Hearing • Taste • Smell • Equilibrium • Somatic Senses
  • 2. Sensory Systems • Somatic sensory • General – transmit impulses from skin, skeletal muscles, and joints • Special senses - hearing, balance, vision • Visceral sensory • Transmit impulses from visceral organs • Special senses - olfaction (smell), gustation (taste)
  • 3. • Stimulus - energy source • Internal • External • Receptors • Sense organs - structures specialized to respond to stimuli • Transducers - stimulus energy converted into action potentials • Conduction • Afferent pathway • Nerve impulses to the CNS • Translation • CNS integration and information processing • Sensation and perception – your reality Properties of Sensory Systems
  • 4. Sensory Pathways • Stimulus as physical energy  sensory receptor acts as a transducer • Stimulus > threshold  action potential to CNS • Integration in CNS  cerebral cortex or acted on subconsciously
  • 5. Classification by Function (Stimuli) • Mechanoreceptors – respond to touch, pressure, vibration, stretch, and itch • Thermoreceptors – sensitive to changes in temperature • Photoreceptors – respond to light energy (e.g., retina) • Chemoreceptors – respond to chemicals (e.g., smell, taste, changes in blood chemistry) • Nociceptors – sensitive to pain-causing stimuli • Osmoreceptors – detect changes in concentration of solutes, osmotic activity • Baroreceptors – detect changes in fluid pressure
  • 6. Classification by Location • Exteroceptors – sensitive to stimuli arising from outside the body • Located at or near body surfaces • Include receptors for touch, pressure, pain, and temperature • Interoceptors – (visceroceptors) receive stimuli from internal viscera • Monitor a variety of stimuli • Proprioceptors – monitor degree of stretch • Located in musculoskeletal organs
  • 8. • General somatic – include touch, pain, vibration, pressure, temperature • Proprioceptive – detect stretch in tendons and muscle provide information on body position, orientation and movement of body in space Somatic Senses
  • 9. Somatic Receptors • Divided into two groups • Free or Unencapsulated nerve endings • Encapsulated nerve endings - consist of one or more neural end fibers enclosed in connective tissue
  • 10. Free Nerve Endings • Abundant in epithelia and underlying connective tissue • Nociceptors - respond to pain • Thermoreceptors - respond to temperature • Two specialized types of free nerve endings • Merkel discs – lie in the epidermis, slowly adapting receptors for light touch • Hair follicle receptors – Rapidly adapting receptors that wrap around hair follicles
  • 11. Encapsulated Nerve Endings • Meissner’s corpuscles • Spiraling nerve ending surrounded by Schwann cells • Occur in the dermal papillae of hairless areas of the skin • Rapidly adapting receptors for discriminative touch • Pacinian corpuscles • Single nerve ending surrounded by layers of flattened Schwann cells • Occur in the hypodermis • Sensitive to deep pressure – rapidly adapting receptors • Ruffini’s corpuscles • Located in the dermis and respond to pressure • Monitor continuous pressure on the skin – adapt slowly
  • 12. Encapsulated Nerve Endings - Proprioceptors • Monitor stretch in locomotory organs • Three types of proprioceptors • Muscle spindles – monitors the changing length of a muscle, imbedded in the perimysium between muscle fascicles • Golgi tendon organs – located near the muscle-tendon junction, monitor tension within tendons • Joint kinesthetic receptors - sensory nerve endings within the joint capsules, sense pressure and position
  • 13. Muscle Spindle & Golgi Tendon Organ
  • 14. Special Senses Figure 10-4: Sensory pathways • Taste, smell, sight, hearing, and balance • Localized – confined to the head region • Receptors are not free endings of sensory neurons but specialized receptor cells
  • 15. Anatomy of the Eyeball • Function of the eyeball • Protect and support the photoreceptors • Gather, focus, and process light into precise images • External walls – composed of three tunics (layers) • Internal cavity – contains fluids (humors)
  • 16. The Fibrous Layer • Most external layer of the eyeball • Cornea • Anterior one-sixth of the fibrous tunic • Composed of stratified Squamous externally, simple squamous internally • Refracts (bends) light • Sclera • Posterior five-sixths of the tunic • White, opaque region composed of dense irregular connective tissue Provides shape and an anchor for eye muscles, • Scleral venous sinus – allows aqueous humor to drain
  • 17. The Vascular Layer • Middle layer consists of choroid, ciliary body, and iris • Iris and Pupil • Composed of smooth muscle, melanocytes, and blood vessels that forms the colored portion of the eye. • Function: It regulates the amount of light entering the eye through the pupil. • It is attached to the ciliary body. • Pupil is the opening in center of iris through which light enters the eye • Ciliary body • Composed of a ring of muscle called ciliary muscle and ciliary processes which are folds located at the posterior surface of ciliary bodies • Suspensory ligaments attach to these processes • Function: secretes the aqueous humor • The suspensory ligaments position the lens so that light passing through the pupil passes through the center of the lens of the eye.
  • 18. The Vascular Layer • Choroid - vascular layer in the wall of the eye. • Dark brown (pigmented) membrane with melanocytes that lines most of the internal surface of the sclera. Has lots of blood vessels • Lines most of the interior of the sclera. • Extends from the ciliary body to the lens. • Corresponds to arachnoid and pia mater • Functions: • Delivers oxygen and nutrients to the retina. • Absorb light rays so that the light rays are not reflected within the eye
  • 19. The Inner Layer (Retina) • Retina is the innermost layer of the eye lining the posterior cavity • The retina contains 2 layers: • Pigmented layer made of a single layer of melanocytes, absorbs light after it passes through the neural layer • Neural layer – sheet of nervous tissue, contains three main types of neurons • Photoreceptor cells • Bipolar cells • Ganglion cells
  • 20. Photoreceptors • Two main types • Rod cells • More sensitive to light • Allow vision in dim light • In periphery • Cone cells • Operate best in bright light • High-acuity • Color vision – blue, green, red cones • Concentrated in fovea
  • 21. Regional Specializations of the Retina • Ora serrata retinae • Neural layer ends at the posterior margin of the ciliary body • Pigmented layer covers ciliary body and posterior surface of the iris • Macula lutea – contains mostly cones • Fovea centralis – contains only cones • Region of highest visual acuity • Optic disc – blind spot
  • 22. The Lens • A thick, transparent, biconvex disc • Held in place by its ciliary zonule • Lens epithelium – covers anterior surface of the lens
  • 23. The Eye as an Optical Device • Structures in the eye bend light rays • Light rays converge on the retina at a single focal point • Light bending structures (refractory media) • The lens, cornea, and humors • Accommodation – curvature of the lens is adjustable • Allows for focusing on nearby objects
  • 24. Internal Chambers and Fluids Figure 16.8
  • 25. Internal Chambers and Fluids • Anterior segment • Divided into anterior and posterior chambers • Anterior chamber – between the cornea and iris • Posterior chamber – between the iris and lens • Filled with aqueous humor • Renewed continuously • Formed as a blood filtrate • Supplies nutrients to the lens and cornea
  • 26. Internal Chambers and Fluids • The lens and ciliary zonules divide the eye • Posterior segment (cavity) • Filled with vitreous humor - clear, jelly-like substance • Transmits light • Supports the posterior surface of the lens • Helps maintain intraocular pressure
  • 27. Accessory Structures of the Eye • Eyebrows – coarse hairs on the superciliary arches • Eyelids (palpebrae) • Separated by the palpebral fissure • Meet at the medial and lateral angles (canthi) • Conjunctiva – transparent mucous membrane • Palpebral conjunctiva • Bulbar (ocular) conjunctiva • Conjunctival sac • Moistens the eye Figure 16.5a
  • 28. Accessory Structures of the Eye • Lacrimal apparatus – keeps the surface of the eye moist • Lacrimal gland – produces lacrimal fluid • Lacrimal sac – fluid empties into nasal cavity Figure 16.5b
  • 29. Extrinsic Eye Muscles Figure 16.6a, b • Six muscles that control movement of the eye • Originate in the walls of the orbit • Insert on outer surface of the eyeball
  • 30. Visual Pathways to the Cerebral Cortex • Pathway begins at the retina • Light activates photoreceptors • Photoreceptors signal bipolar cells • Bipolar cells signal ganglion cells • Axons of ganglion cells exit eye as the optic nerve
  • 31. • Optic nerve • Optic chiasm • Optic tract • Thalamus • Visual cortex • Other pathways include the midbrain and diencephalon Vision Integration / Pathway Figure 10-29b, c: Neural pathways for vision and the papillary reflex
  • 32. The Ear: Hearing and Equilibrium • The ear – receptor organ for hearing and equilibrium • Composed of three main regions • Outer ear – functions in hearing • Middle ear – functions in hearing • Inner ear – functions in both hearing and equilibrium
  • 33. The Outer (External) Ear • Auricle (pinna) - helps direct sounds • External acoustic meatus • Lined with skin • Contains hairs, sebaceous glands, and ceruminous glands • Tympanic membrane • Forms the boundary between the external and middle ear
  • 34. The Middle Ear • The tympanic cavity • A small, air-filled space • Located within the petrous portion of the temporal bone • Medial wall is penetrated by • Oval window • Round window • Pharyngotympanic tube (auditory or eustachian tube) - Links the middle ear and pharynx
  • 35. Figure 16.17 The Middle Ear • Ear ossicles – smallest bones in the body • Malleus – attaches to the eardrum • Incus – between the malleus and stapes • Stapes – vibrates against the oval window
  • 36. The Inner (Internal) Ear • Inner ear – also called the labyrinth • Bony labyrinth – a cavity consisting of three parts • Semicircular canals • Vestibule • Cochlea • Bony labyrinth is filled with perilymph
  • 37. The Membranous Labyrinth Figure 16.18 • Membranous labyrinth - series of membrane-walled sacs and ducts • Fit within the bony labyrinth • Consists of three main parts • Semicircular ducts • Utricle and saccule • Cochlear duct • Filled with a clear fluid – endolymph
  • 38. The Cochlea • A spiraling chamber in the bony labyrinth • Coils around a pillar of bone – the modiolus • Spiral lamina – a spiral of bone in the modiolus • The cochlear nerve runs through the core of the modiolus
  • 39. The Cochlea • The cochlear duct (scala media) – contains receptors for hearing • Lies between two chambers • The scala vestibuli • The scala tympani • The vestibular membrane – the roof of the cochlear duct • The basilar membrane – the floor of the cochlear duct
  • 40. The Cochlea • The cochlear duct (scala media) – contains receptors for hearing • Organ of Corti – the receptor epithelium for hearing • Consists of hair cells (receptor cells)
  • 41. The Role of the Cochlea in Hearing Figure 16.20
  • 42. Auditory Pathway from the Organ of Corti • The ascending auditory pathway • Transmits information from cochlear receptors to the cerebral cortex Figure 16.23
  • 43. The Vestibule • Utricle and saccule – suspended in perilymph • Two egg-shaped parts of the membranous labyrinth • House the macula – a spot of sensory epithelium • Macula – contains receptor cells • Monitor the position of the head when the head is still • Contains columnar supporting cells • Receptor cells – called hair cells • Synapse with the vestibular nerve
  • 44. Anatomy and Function of the Maculae Figure 16.21b
  • 45. The Semicircular Canals • Lie posterior and lateral to the vestibule • Anterior and posterior semicircular canals lie in the vertical plane at right angles • Lateral semicircular canal lies in the horizontal plane
  • 46. The Semicircular Canals • Semicircular duct – snakes through each semicircular canal • Membranous ampulla – located within bony ampulla • Houses a structure called a crista ampullaris • Cristae contain receptor cells of rotational acceleration • Epithelium contains supporting cells and receptor hair cells
  • 47. Structure and Function of the Crista Ampullaris Figure 16.22b
  • 48. The Chemical Senses: Taste and Smell • Taste – gustation • Smell – olfaction • Receptors – classified as chemoreceptors • Respond to chemicals
  • 49. Taste – Gustation • Taste receptors • Occur in taste buds • Most are found on the surface of the tongue • Located within tongue papillae • Two types of papillae (with taste buds) • Fungiform papillae • Circumvallate papillae
  • 50. Taste Buds • Collection of 50 –100 epithelial cells • Contain three major cell types (similar in all special senses) • Supporting cells • Gustatory cells • Basal cells • Contain long microvilli – extend through a taste pore
  • 51. Taste Sensation and the Gustatory Pathway • Four basic qualities of taste • Sweet, sour, salty, and bitter • A fifth taste – umami, “deliciousness” • No structural difference among taste buds
  • 52. Gustatory Pathway from Taste Buds Figure 16.2 • Taste information reaches the cerebral cortex • Primarily through the facial (VII) and glossopharyngeal (IX) nerves • Some taste information through the vagus nerve (X) • Sensory neurons synapse in the medulla • Located in the solitary nucleus
  • 53. • Olfactory epithelium with olfactory receptors, supporting cells, basal cells • Olfactory receptors are modified neurons • Surfaces are coated with secretions from olfactory glands • Olfactory reception involves detecting dissolved chemicals as they interact with odorant binding proteins Smell (Olfaction)
  • 54. Olfactory Receptors • Bipolar sensory neurons located within olfactory epithelium • Dendrite projects into nasal cavity, terminates in cilia • Axon projects directly up into olfactory bulb of cerebrum • Olfactory bulb projects to olfactory cortex, hippocampus, and amygdaloid nuclei