3. 1. Discuss the three distribution patterns
of individuals within a population and
explain the condition that give rise to
each pattern.
4. 3 Distribution Patterns of Individuals
Source: https://www.boundless.com/biology/population-and-community-
ecology/population-demography/species-distribution
5. Random- individuals are distributed
randomly if each individual position is
independent of those of the others.
Plants that have wind-dispersed
seeds such as dandelions exhibit
random distribution as they
germinate wherever they happen to
fall in a favorable environment.
Source: https://www.boundless.com/biology/population-and-community-
ecology/population-demography/species-distribution
6. Uniform- individuals are more or less
evenly spaced. A uniform distribution
usually results from some form of
negative interaction among individuals
such as competition, which functions to
maintain minimum distance among
members of the population.
7. In animals- uniform distribution is
common where individuals defend an
area for their exclusive use
(territoriality)
In plants- uniform distribution is
common where severe competition
exists for belowground resources such
as water or nutrients
Source: Elements of Ecology, 8th Edition
8. Clump- most common distribution
which individuals occur in groups.
Clumping of individuals happen when
suitable habitat or other resources
may be distributed as patches on the
larger landscape, forming of social
groups among individuals and asexual
reproduction
Source: Elements of Ecology, 8th Edition
9. 2. Provide the mark-recapture
equation used for estimating
population size and define the
terms of the equation.Why is the
mark-recapture such a popular &
effective method for determining
animal population densities?
10. MARK-RECAPTURE EQUATION
- A method based on trapping,
marking, and releasing a known
number of marked animals into
the population.
*This procedure was first used by C.J.G.
Petersen in studies of marine fishes & F.C
Lincoln in studies of waterfowl populations.
12. This technique is commonly used by
fish and wildlife managers to estimate
population sizes before fishing or
hunting seasons. Ecologists also find
that a measure of relative density or
abundance is sufficient and by using
mark-recapture method trends in
abundance is depicted thus this
method is popular and effective.
Source: http://www.radford.edu
13. 3. Describe several approaches
used by ecologists to establish age
structure for plant and animal
populations
14. Some approaches used by ecologists
to establish age structure
Animals:
âą Marking young individuals in a population
& following their survival through time
âą Examining sample of individual carcasses
to determine age at death
âą Morphological changes such as wear &
replacement of teeth among deers or
counting rings deposited annually in fish
15. Some approaches used by ecologists
to establish age structure
Plants:
âą Marking young individuals in a population
& following their survival through time
âą Determining size of diameter (assumption
that diameter increases with age)
âą Attempts to age non-woody plants have
not been successful
Source: Elements of Ecology, 8th Edition
16. 4. Some animals migrate daily
while others migrate seasonally.
Give an example of each migration
pattern and explain the benefit of
this lifestyle to the organisms that
you present.
17.
18. 5. Using examples, discuss the
impact of increased human
travel in the 20th century on
long-distance dispersal of
plants & animals. Describe how
& why introduced species
typically alter native
ecosystems
19. In human-dominated landscapes, people have
a major influence not only on the distribution,
but also on the dispersal of plant & animal
species . Human-mediated dispersal in these
environments can be either intentional or
unintentional-A. Auffret & S. Cousins
(Humans as Long-Distance Dispersers of
Rural Plant Communities)
20. Examples of Human assisted dispersal of species
1. Dispersal of seeds on human clothes and
footwear
2. Seeds of weeds included in shipments of
imported crops
3. Domestic pet trade such as Janitor fish
4. Introduction of new species as raw materials
5. Domestic trade of ornamental plants
21. Though not all human-dispersed species can
create imbalance in any native ecosystem, those
that become invasive species do endanger the
species living in the native ecosystem.
The direct threats of invasive species are:
1.Preying on native species
2.Out-competing native species for food or other
resources
3.Causing or carrying disease
4.Preventing native species from reproducing or
22. The indirect threats of invasive species are:
1.Changing food webs
2.Decreasing biodiversity
3.Altering ecosystem conditions
Source: http://www.nwf.org
24. 1.What is the significance of
constructing a life table to an
ecologist?What information is
required to develop a life table and
what is this information used for?
25. What is a LifeTable?
* A more sophisticated method for
examining population abundance is to
construct a life table
*This table will have a schedule of all
births and deaths in all, or more likely
some portion, of a population
26. What information can you get
from a life table?
*Population age structure âAre there lots of:
young individuals? Old individuals?
Reproductive age individuals
* Population growth rate âHow fast is the
population size growing (or shrinking)?
* Population survivorship patterns âDoes most
mortality occur in the very young?The very old?
Or equally across all ages?
Source: http://iweb.tntech.edu
32. Type 1:
Humans:
low mortality rate
(early on) and give
birth to few
young. Parental
care is high, rarely
preyed on, and
migration is low,
thus, more likely
to survive.
Type 2:
Squirrel:
Mortality rate is
common (linear),
birth rate is
medium to high,
and migrate an
average amount.
Parental care is
also not as long as
Type 1 andType
2s are more
preyed on.
Type 3:
Clam:
Birth rate is
extremely high
and mortality rate
is extremely high.
There is little to
usually no
parental care.
Source: www.uic.edu
33. 4. Explain how λ is used by
demographers to predict the future
size of a population.
34. Lambda (λ) is called the finite population
growth rate that gives the proportional
change in population size from one time
period to the next:
Where:
đ”đ + đ đąđŹ đđšđđđ„ đ§đźđŠđđđ« đšđ đąđ§đđąđŻđąđđźđđ„đŹ đąđ§ đČđđđ« đ + đ
Nt is the total number of individuals in the previous year
Source: Elements of Ecology, 8th Ed.
36. 5. Discuss 5 factors known to
cause extinction.Which of these
factors is considered the leading
cause of current extinctions and
why?
37. 5 Factors known to cause extinction:
1. Extreme environmental events
2. Overexploitation of resources
3. Invasive species in a native ecosystem
4. Loss of habitat due to human activities
5. Disease
38. Developmental activities such as urbanization, agriculture,
mining and deforestation had greatly affected the habitats of
many species. Many of which were forced to look for a more
suitable niche which could also mean that those said organisms
would have to adapt again to a new environment. Adaptation in
this case can be positive or can be detrimental to the organisms.
The latter could cause their extinction.
39. Source: Quantifying threats to imperiled species in the United States
According to a study conducted in 1998 by Wilcove,
Rothstein, Dubow, Phillips, and Losos (Quantifying threats
to imperiled species in the United States), the number one
cause of endangerment of species is extinction due to
human activities.
40. 6.What is the Allee effect? Give
several examples that illustrate
the potential catastrophic
impact of small size on
population persistence.
41. Allee Effect- The decline in either
reproduction or survival under
conditions of low population density
(Smith & Smith, 2012)
42. Examples illustrating Allee Effect:
1. Difficulty in finding mate for large cats
that are widely spread
2. Species using chemicals such as
pheromones to communicate and
attract mates
3. Likeliness of plants being pollinated
4. Predation on island fox (Urocyon
littoralis) by Golden Eagles
Sources: Elements of Ecology, 8th Ed.
http://www.ncbi.nlm.nih.gov
46. Sources: S. Otto, 2008- Sexual reproduction and the evolution of sex
http://www.ck12.org
47. 2. Explain the advantage of the
ability for an organism to possess
both male and female organs
48. An organism that possess both a female and a male
Sexual organ is called a hermaphrodite.
Hermaphroditism is quite common in invertebrates
& plants and exceedingly rare in vertebrates.
Most Perciforms (fish) are naturally hermaphrodite.
50. Advantages:
1. Low density model
2. Size advantage model
3. Gene dispersal model
Source: M. Ghiselin âThe Evolution of Hermaphroditism Among Animals (1969)
51. 3. Explain why most bird species
are monogamous while most
mammal species are polygamous.
52. âIn all sexually reproducing species
there is a social framework involving
the selection of mates.The pattern of
mating between males and females in
a population is called mating
systemâ- Smith & Smith, 2012
53. Monogamy is common among birds because
most young birds are helpless at hatching
and need food, warmth and protection.
Male birds can increase his fitness more by
continuing his investment in the young.Also,
parental care of both parents is necessary to
raise the young successfully. Parents
alternate in giving protection and
nourishment to the young birds.
54. In mammals, polygamy is a common
practice. It can involve 1 male and several
female or vice versa.The individual having
multiple mates is generally not involved in
caring for the young. For some, since females
lactate, providing food for the young it is to
the advantage of the males to mate with as
many females as possible.
Source: Elements of Ecology, 8th. Ed.
56. Darwin considered that most sexual dimorphism
was due to sexual selection, in which evolutionary
forces acted separately on the sexes.
Source: www.bbc.co.ukSource: www.bbc.co.uk
Source: http://www.cetus.ucsd.edu
57. 5. Contrast semelparous and
iteroparous reproductive
strategies and explain the
trade-offs of each.
59. 6. Explain why siblicide is common
among bird species when resources
are rare.Why is this an effective
strategy under these conditions?
60. Characteristics common to all siblicide:
1. Competition for food
2. Provision of food to the nestlings in small units
3. Weaponry
4. Competitive disparities between siblings
5. Spatial confinement Source: Avian Siblicide âR. Santana, D. Mock, H.
Drummond, H. Stinson
61. Parental Favoritism and Siblicide
Sometimes parents seem not only to tolerate
siblicide, but to actively encourage it.
For example, in Black Eagles incubation begins as
soon as the first egg is laid.
As a result the first egg hatches 3-7 days before the
second and so the older offspring has a huge size
advantage over its younger sibling and can easily
kill him.
62. Why parents play favorites and facilitate
siblicide?
There are two major reasons:
* Insurance against failure
* Environmental uncertainty
63. 7. Discuss the differences between
r-strategists and K- strategists.
Give an example of each.
67. Ruderals (R)
âą rapidly colonize disturbed sites
âą small in stature and short lived
âą allocation of resources is primarily to
reproduction with characteristics allowing for a
wide dispersal of seeds to newly disturbed
sites
68. Competitive (C)
âą thrive in predictable habitats with abundant
resources
âą Allocate resources to growth favoring resource
acquisition and competitive ability
Stress-Tolerant (S)
âą Thrive in areas where resources are limited
âą Allocate resource to maintenance
70. 1.What is the difference
between exponential growth
and logistic growth? Under
what ecological conditions does
each occur?
71. Exponential Population Growth-
characteristic of populations inhabiting
favorable environments at low densities such
as during the process of colonization &
establishment in new environments- Smith &
Smith (2012)
Associated with the name ofThomas Robert
Malthus (1766-1834) who first realized that
any species can potentially increase in
numbers according to a geometric series.
Source: A. Sharov, Quantitative Population Ecology
72. 3 possible model outcomes:
Source: A. Sharov, Quantitative Population Ecology
73. Assumptions of Exponential Model:
1. Continuous reproduction (e.g., no seasonality)
2. All organisms are identical (e.g., no age
structure)
3. Environment is constant in space and time
(e.g., resources are unlimited)
Source: A. Sharov, Quantitative Population Ecology
74. Logistic population growth- developed by
Belgian mathematician PierreVerhulst
(1838) who suggested that the rate of
population increase may be limited, i.e., it
may depend on population density
75. 3 possible model outcomes:
Logistic model combines two ecological processes: reproduction
and competition. Both processes depend on population numbers
(or density).The rate of both processes corresponds to the mass-
action law with coefficients: r0 for reproduction and ro/K for
competition.
76. 2. Discuss some of the
important issues involved when
trying to determine Earthâs
carrying capacity (K) for
humans.Why is it difficult to
calculate K for the human
population?
77. Scientists have been in the quest to find the
exact carrying capacity of Earth among human
beings. Aside from answering the question âhow
manyâ, scientists have also quantified the
questions to â at what standard of living & at
what cost to the environment is the human
carrying capacity of the Earth has?
78. 2 scientist who presented their estimate of the
Earthâs carrying capacity (K) on humans:
1.C.T. DeWit- based solely on estimates of primary
production and the average annual per capita energy
requirement estimated that 1000 billion people could
live from Earth if photosynthesis is the limiting factor
2.H. R. Hulett- reached a conclusion that approximately
1 billion people represented the maximum world
population supportable at the then current agricultural
& industrial levels of production.
Source: Elements of Ecology, 8th Ed.
79. Issues about the conducted studies:
DeWit- uses assumptions in his calculation of
the Earthâs carrying capacity such as sufficient
water and nutrients for all human beings and an
assumption that the entire land area is
dedicated to the production of plants.
Hulett- used the standard of living and resource
consumption for the average U.S citizen as his
definition of optimal lifestyle.
80. 3. Contrast the density-dependent and
density-independent mechanism of
population regulation.
83. Competition- âcompetition is an interaction
between individuals, brought about by a shared
requirement for a resource [in limited supply],
and leading to a reduction in the survivorship,
growth and/or reproduction of at least some of
the competing individuals concernedâ
(Begon et al., 2006)
Intraspecific competition- competition among
individuals of the same species
85. 5. Explain why high population density
might stress an organism. Provide
several examples of some of the
negative impacts of stress on an
organism.
86. High density population is stressful to individuals
since restricted space can often lead to aggressive
contacts among individuals. Increased crowding
and social contact can cause stress.
Stress triggers:
1. Hormonal changes
2. Suppress immune system
3. Increase mortality of the young in the fetal
stage (decreased birth & increased mortality)
87. 6.Why are social organization and
territoriality adaptive? How do
human populations exhibit these
two behaviors?
88. Social behavior appears to be a mechanism that
limits the number of animals living in a particular
habitat, having access to a common food supply,
and engaging in reproductive activities.
Social dominance plays a role in population
regulation when it affects reproduction &
survival in a density-dependent manner.
Territoriality secures the individual sole access
to an area of habitat and the resources it
contains.
89. 7.Why do some species have a
larger home range than others?
Explain why a carnivorous
mammal requires a larger range
than an herbivorous mammal of
similar body size?
90. Home range- the area that an animal normally
uses during a year.
Overall size of the home range varies with the
available food resources, mode of food
gathering, body size, and metabolic needs.
91. For a given body mass, the home range of
carnivores is larger than that of herbivores
because the home range of a carnivore must be
large enough to support a population of the
prey that it feeds upon- Harastad & Bunnell, 1979
Hinweis der Redaktion
Methods involve observations relating to the presence of organisms rather than to direct counts of the individuals. If these observations have some relatively constant relationship to total individuals seen per kilometer or heard per hour, such count called indices of abundance and if collected from the same area over a period of years, it depicts trends in abundance.
The indirect threats of invasive species:
Changing food webs: Invasive species can change the food web in an ecosystem by destroying or replacing native food sources. The invasive species may provide little to no food value for wildlife.
Decreasing biodiversity: Invasive species can alter the abundance or diversity of species that are important habitat for native wildlife. Aggressive plant species like kudzu can quickly replace a diverse ecosystem with a monoculture of just kudzu.
Altering ecosystem conditions: Some invasive species are capable of changing the conditions in an ecosystem, such as changing soil chemistry or the intensity of wildfires.
Cohort- a group of individuals of the same age
Small populations can be susceptible to a variety of factors that directly influence the rates of survival and birth. If only a few individuals make up a population, the fate of each individual can be crucial to the survival of the population.
Sex Evolves When Selection Changes Over Time: Current models indicate that sex evolves more readily when a species' environment changes rapidly. When the genetic associations built up by past selection are no longer favorable, sex and recombination can improve the fitness of offspring, thereby turning the recombination load into an advantage.Â
Sex Evolves When Organisms Are Less Adapted to Their Environment: Organisms that reproduce both sexually and asexually tend to switch to sex under stressful conditions. Mathematical models have revealed that it is much easier for sex to evolve if individuals that are adapted to their environment reproduce asexually and less fit individuals reproduce sexually.
Low density model- invokes the advantage of being able to mate with any other member of the species where the probability of encountering a suitable mate is low.
Size advantage model- explains sequential hermaphroditism as occurring where an individual reproduces most efficiently as a member of one sex when small or young, but as a member of the other sex when it gets older or
larger; it predicts proterogyny where there is sexual selection for larger males, and protandry where the young stages must hunt for a suitable
Gene dispersal model- is based upon the idea that the individual may be adapted to the genetical environment, or to the population structure as affected by the prevailing conditions of gene flow.
Among the mammalian exceptions, the male provides for the female and young and defends the territory (area defended for exclusive use and access to resources. In addition, freed from parental duty, the individual can devote more time and energy to competition for more mates and resources.
For example, females might choose to mate with highly ornamented males (e.g., the peacockâs tail) or males might develop characters useful for ïŹghting with other males to win in contests for access to females (e.g., large body size and antlers in deer). Today, these two processes are often referred to as female choice and contest competition, respectively.
Some cases of sexual dimorphism seem to be best explained by natural selection: For example, males and females of in some species of birds have radically different bill morphologies that are best explained by sex differences in foraging habits (Andersson, 1994).
Later reproduction means increased growth later maturity and increased survivorship but less time for reproduction.
What type of change in conditions might bring about the shift from semelparity to iteroparity?
If the external environment imposes a high adult mortality relative to juvenile mortality and if individuals reach maturity, chances are that they will not survive much longer so future reproductive expectations are bleak. Semelparity is favored.
If the opposite holds true and juvenile mortality is high compared to adult mortality, once an individual makes it to maturity, it has a good chance of surviving into the future so prospects of future reproduction are good. Iteroparity is favored.
Shortage of food
Ability to defend each unit of food
Possess lethal attack against the siblings
Large compared to small or the weaklings
Competition for space in the nest
Such hatching asynchrony is very common among birds and results in a hatching asynchrony that establishes an age and size hierarchy within the brood.
In all siblicidal species studied to date there is a striking tendency for the victim to be the youngest member of the brood (Mock and Parker 1986). The youngest sibling is marginal in the sense that its reproductive value can be assessed in terms of what it adds to or subtracts from the success of other family members. Specifically, the marginal
Individual can embody two kinds of reproductive value. First, if the marginal individual survives in addition to all its siblings, it represents an extra unit of parental success, or extra reproductive value. Such an event is most likely during an especially favorable season, when the needs of the entire brood can be satisfied. Alternatively, the marginal offspring may serve as a replacement for an elder sibling that dies prematurely.
In such instances the marginal individual represents a form of insurance against the loss of a senior sibling.
The magnitude of this insurance value depends on the probability that the senior sibling will die.
These animals have no intention of rearing more than a single offspring.
The second offspring represents an easily cancelled insurance policy against the failure of the first offspring to hatch or develop normally.
When the first offspring arrives it kills its sibling (Black Eagle), the parents cover over the second egg (Harpy Eagles), the parents abandon the second egg (Hooded Grebes) or abandon the second born cub (Giant Pandas).
Thus, the parents avoid prolonged investment in a back-up offspring. However, if the first offspring fails the second can step in and take its place.
If food is plentiful, the younger chicks can tolerate the beating and may survive to fledge. If food is scarce the younger chicks quickly starve or are driven out of the nest and die.
K-selection and r-selection are the two broad categories of life history strategies. The life history strategy of a species incorporates aspects of how the organism reproduces, its strategies for survival, how the organism interacts with its habitat, and how it is able to compete with other organisms within the habitat.
These characteristics describe r-selected species perfectly:
Rapid Development
High Reproductive Rate
Early Reproductive Age
Small Body Size
One Reproductive Cycle
Short Lifespan
Poor Competitive Ability
High Mortality of Offspring
Population Below the Carrying Capacity of the Habitat
Good Offspring Dispersal
Found in Disturbed Habitats
Limited or No Parental Care
The following is a list of key K-selected traits:
Slow Development
Low Reproductive Rate
Late Reproductive Age
Large Body Size
Multiple Reproductive Cycles
Long Lifespan
Strong Competitive Ability
Low Offspring Mortality
Relatively Steady Population Size near the Carrying Capacity
Limited Offspring Dispersal
Found in Mature, Undisturbed Habitats
Good Parental Care
However, exponential model is robust; it gives reasonable precision even if these conditions do not met. Organisms may differ in their age, survival, and mortality. But the population consists of a large number of organisms, and thus their birth and death rates are averaged.
Parameter ro is relatively easy to interpret: this is the maximum possible rate of population growth which is the net effect of reproduction and mortality (excluding density-dependent mortality). Slowly reproducing organisms (elephants) have low ro and rapidly reproducing organisms (majority of pest insects) have high ro. The problem with the logistic model is that parameter ro controls not only population growth rate, but population decline rate (at N > K) as well. Here biological sense becomes not clear. It is not obvious that organisms with a low reproduction rate should die at the same slow rate. If reproduction is slow and mortality is fast, then the logistic model will not work.
Parameter KÂ has biological meaning for populations with a strong interaction among individuals that controls their reproduction. For example, rodents have social structure that controls reproduction, birds have territoriality, plants compete for space and light. However, parameter K has no clear meaning for organisms whose population dynamics is determined by the balance of reproduction and mortality processes (e.g., most insect populations). In this case the equilibrium population density does not necessary correspond to the amount of resources; thus, the term "carrying capacity" becomes confusing. For example, equilibrium density may depend on mortality caused by natural enemies.
DeWit- uses assumptions in his calculation of the Earthâs carrying capacity such as sufficient water and nutrients for all human beings and an assumption that the entire land area is dedicated to the production of plants.
Hulett- used the standard of living and resource consumption for the average U.S citizen as his definition of optimal lifestyle.
DeWit- uses assumptions in his calculation of the Earthâs carrying capacity such as sufficient water and nutrients for all human beings and an assumption that the entire land area is dedicated to the production of plants.
Hulett- used the standard of living and resource consumption for the average U.S citizen as his definition of optimal lifestyle.
Exploitation- competing individuals do not directly interact with one another instead respond to the level of resource availability that is depressed by the presence and consumption of other individuals in the population
Interference- individuals interact directly with each other, preventing others from occupying a habitat accessing resources within it.
Hormonal changes- can suppress growth, curtail reproductive functions & delay sexual activity
Suppress immune system- break down the white blood cells, increasing vulnerability to disease
Humans have many similarities to animal's territorial behavior, only in a more sophisticated way (I don't believe you wake up every morning to pee around your lawn to mark it as yours, do you?).
It starts at the grand scale â countries, each with its own borders and armies to guard it. Then we can scale down to: regions, cites, streets, your neighborhood, your house, your room, your table and finally â your favorite mug!
Just like other animals, we like to border our stuff and mark them as ours in various ways (more on that later). There is also a strong link between social status and the ownership and size of territory, the bigger it is - the higher the status (in most cases).
On the flip side, we display some distinct behaviors that separate us from other animals:
First of all, we use territory for a variety of purposes besides breeding and securing food resources. We have special places for: work, recreation, sports, study or just hanging out. We also guard our territory out of different reasons; if you find your house robbed you won't check the fridge to look for missing food!
Secondly, it's not very common to act aggressively towards others who invade your space. You might get angry yes, but it doesn't mean that you'll bare your teeth and prepare for an attack. Moreover, we often act oppositely to territoriality by inviting others to our place for social or technical purposes.
And lastly, we don't mark our territory with our pee (:
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