3. 1. Describe the effects of temperature,
moisture, length of photosynthetic period,
and nutrient availability on net primary
productivity in terrestrial ecosystems.
4. Terms to define:
Net Primary Productivity (NPP)- is the rate of
energy storage as organic matter after respiration
Gross Primary Productivity (GPP)- total rate of
photosynthesis or energy assimilated by the
autothrophs
5. Factors limiting primary productivity in terrestrial
ecosystems
Temperature
Precipitation
Light
Nutrients
9. To summarize:
Temperature + Precipitation = NPP
Temperature + Precipitation = NPP
Temperature + Precipitation = NPP
Temperature = Length of Photosynthesis = NPP
Nutrients = Nutrient uptake,
photosynthesis & plant growth =
NPP
10. 2. Explain why primary
productivity is typically higher in
coastal areas of the ocean than far
from shore in the open ocean.
11.
12. 3. Explain the difference between
primary production and secondary
production. How is secondary
production affected by primary
production?
13. Primary Production- production by
green plants over time
Secondary Production- production by
consumer organisms over time which
include consumption of energy from
the produces which is then used for
metabolic activities
15. 4. What are the two main food chains
of ecosystems? Which is more
dominant in terrestrial ecosystems and
why?
16. Grazing food chain- the source of energy is living
plant biomass or net primary production
ex. Cattle grazing on pastureland, insect feeding
on leaves, zooplankton feeding on phytoplankton
Detrital food chain- source of energy is dead
organic matter or detritus
ex. snails, bettles, millipeds, earthworms, fungi &
bacteria
18. Detrital food chain is the most common food
chain in all ecosystems since most ungrazed or
unconsumed energy is returned to the soil as
dead material.
In a yellow poplar forest
maintenance & respiration 50%
new tissue 13%
consumed by herbivores 2%
Detrital food chain 35%
19. 5. Explain why energy is lost as it is
transferred from one trophic level to
another. What are the effects of energy
loss on the biomass of different trophic
levels?
22. 6. How do humans affect net primary
productivity? Does the affluence of a
society affect how humans appropriate
net primary productivity?
23. • Human species consume a vast amount of
Earth’s resources
• In a study conducted by Vitousek, et. al they
used 3 approaches to estimate human NPP
appropriation:
low estimate- food, fuel, fiber, timber
intermediate estimate- productivity of
lands devoted to human activities
high estimate- conversion of lands to
cities, forests to pastures or overgrazing
or excessive erosion
24. Both population and per capita consumption
interact to determine the human ecological
impact on regional scale.
Affluence also affect net primary productivity
among humans. Developed countries have
double HANPP compared to developing nations
26. 1. Describe the movement of an atom of
nitrogen from the leaf of a plant,
through the process of decomposition,
and back into the root of another plant.
27.
28.
29. 2. Describe three different processes
that are important for decomposition.
Explain which organisms are important
for each of the processes.
30. Decomposition processes:
1. Leaching- dissolving and washing of
nutrients out of soil, litter and organic matter
ex. bacteria and fungi
2. Fragmentation- disintegration of organic
matter by invertebrate detritivores such as
protozoans, nematodes, millipedes, etc.
3. Ingestion & excretion of waste products-
heterotrophs function as decomposers by
digesting food, break down organic matter,
31. 3. Discuss the effects of temperature,
humidity, and oxygen availability on
the rate of decomposition.
32. 2 things related to decomposition rate:
the quality of plant litter as a substrate for
microorganisms and soil fauna active in the
decomposition process
Features of the physical environment that
directly influence decomposer populations
like soil properties and climate
33.
34.
35. 4. Explain how the "quality" of plant-
derived organic matter changes during
the course of decomposition.
38. Mineralization- microbial breakdown of humus and
other organic matter in soil to inorganic matter esp
into its ionic form such as ammonium, sulfate and
phosphate
Immobilization- conversion of an element from
inorganic to organic form in microbial or plant tissue,
rendering the nutrient unavailable to other organisms
39.
40. 6. Describe two major differences between
decomposition in terrestrial environments
and
decomposition in aquatic environments.
41. Terrestrial Decomposition
decomposition is a soil process
involves two size-based animal groups; the
macrofauna and the microfauna
primary production and decomposition is linked
by vegetation
(Source: http://ww2.tnstate.edu)
42. Aquatic Decomposition
macrofauna found in freshwater is similar to
those found in soils except for millipedes, in
marine, there are no insects instead there are
polychaetes and many crustaceans
in aquatic systems, the photic zone may be
separated from the benthos (where most
decomposition goes on)
nutrients are only mixed into the photic zone
when thermocline breaks down and Turnover
of the body of water can occur
(Source: http://ww2.tnstate.edu)
43.
44. 7. Describe one major difference between
nutrient cycling in a river and nutrient
cycling in a lake
47. 8. Describe the process of upwelling in
the ocean and how it relates to fisheries
48. What is upwelling?
Winds blowing across the
ocean surface push water
away. Water then rises up
from beneath the surface to
replace the water that was
pushed away
Source: www.oceanservice.noaa.gov
49. Water that rises to the surface as a result of
upwelling is rich in nutrients. These nutrients
“fertilize” surface waters and therefore are good
fishing grounds due too high biological
productivity
50. 9. Describe how you might go about
measuring the rate of decomposition of leaf
litter in the upper layer of a forest soil.
52. 1. Explain the difference between the
two basic types of biogeochemical
cycles, using specific examples of each.
53. What is biogeochemical cycle?
•any of the natural circulation pathways of the
essential elements of living matter. These elements
in various forms flow from the nonliving (abiotic)
to the living (biotic) components of the biosphere
and back to the nonliving again
54. 2 Major types of biogeochemical cycles:
1. gaseous- main pool of nutrients are the
atmosphere and oceans (via evaporation)ex.
carbon, nitrogen, oxgen, carbon and water
2. sedimentary- main pool of nutrients are the
rocks, soil and minerals (Earth’s crust) ex. iron,
phosphorous, sulfur & other earthbound
elements
56. 3 ways of nutrient loss in the ecosystem:
1. Export of nutrients
carbon being exported to the atmosphere via
respiration
microbial and plant processes being transformed
to nutrients in a gaseous phase
Organic matter being carried from the ecosystem
through surface flow of water in streams & rivers
57. 3 ways of nutrient loss in the ecosystem:
2. Harvesting
Biomass harvest such as in farming and logging
3. Fire
Kills vegetation and converts some biomass to soil
organic matter to ash
If not taken up by vegetation during recovery,
nutrients may be lost through leaching and
erosion
58. 3. Describe how atmospheric carbon dioxide
levels fluctuate daily, seasonally, and
geographically. Explain why such fluctuation
occurs.
59. Atmospheric carbon dioxide levels fluctuate on a
daily basis due to:
consistency
photosynthetic stages during the day
less carbon dioxide gathered during sunlight
gathering
60. Atmospheric carbon dioxide levels fluctuate
seasonally and geographically due to:
temperature
solar radiation
timing of the growing and dormant seasons
land area
61. 4. Describe a pathway of nitrogen from the
atmosphere into the body tissue of an
herbivore.
63. 5. Describe how an atom of phosphorus
might move from the crystal lattice of a
mineral in a rock, through a plant, and into
the ocean.
64. phosphate salts are release from rocks
through weathering, leaching, erosion,
mining or agricultural fertilizers usually
dissolved in water
dissolved phosphate being absorbed by
plants
animals absorb phosphates by eating plants
or plant-eating animals
when animals or plants die, phosphates
returns to the soil or ocean
(source: http://www.lenntech.com)
65. 6. List and describe three different inputs of
sulfur into the atmosphere.
66. Sulfur enters the atmosphere from several sources:
1. combustion of fossil fuels
2. volcanic eruptions
3. exchange at the surface of the oceans
67. 7. Describe how ozone is formed at different
levels of the atmosphere and how biological
organisms are affected by ozone.
A variety of environmental factors (climate, radiation) influence productivity
Mean annual temperature: higher temp also means longer growing season - both higher rates and more days over which there is photosynthesis
The higher the rainfall more water available for transpiration, stoma can remain open and photosynthesis can take place longer.
Interaction of temperature and moisture; warmer temperature increase evapotranspiration, there increase water demand if not enough water, productivity will be low
Need both high temperature and high moisture for highest production
In addition to climate, the availability of essential nutrients required for plant growth has a direct effect on ecosystem productivity by influencing nutrient uptake, photosynthesis and plant growth
John Pastor, U of Min in Blackhawk Island, Wisconsin, clearly show relationship between nitrogen availability and aboveground primary productivity
Most productive waters of the ocean are the shallow waters of the coastal environments: 1 shallow waters allow for greater transport of nutrients from bottom sediments to surface waters than deeper waters. (2) coastal regions receive a lot of inputs from terrestrial ecosystems 3. since light is a limiting productivity factor in aquatic ecosystems, open oceans generally have deeper depths thus photosynthetic rate declines as light intensity decreases.
Primary production is a limiting factor to secondary production. Secondary production depends on primary production for energy
Herbivore biomass and consumption of primary productivity by herbivores increase with primary productivity
Orange arrows linking trophic levels represent the flow of energy associated with ingestion. The blue arrows from each trophic level represent the loss of energy through respiration. The brown arrows represent the combination of dead organic matter (unconsumed biomass) and waste products (feces and urine)
In most terrestrial and shallow-water ecosystems, with high standing biomass and relatively low harvest of primary production by herbivores, the detrital food chain is dominant. In deep-water aquatic ecosystems, with their low standing biomass, rapid turnover of organisms, and high rate of harvest, the grazing food chain may be dominant.
Energy decreases in each successive trophic level
Ecological rule of thumb is that only 10 percent of the energy stored as biomass in a given trophic level is converted to biomass at the next higher trophic level
Trophic efficiency - ratio of productivity in a given trophic level with the trophic level upon which it feeds P/P, measure of efficiency that is used to describe he transfer of energy between trophic levels
Important consequence of decreasing energy transfers through the food web is a corresponding decrease in the standing biomass of organisms within each successive trophic level
Exception in some lakes and seas
Most essential nutrients are recycled within the ecosystem
Plant uptake of nutrients in mineral or inorganic form,
Incorporation (nitrogen into enzymes, proteins)
Retranslocation (from leaves about to be shed) up to 70% can be recovered
Bacteria and fungi secrete enzymes into plant and animal tissues to break down the complex organic compounds
Fragmentation of leaves, twigs and other organic matter.
Heterotrophs function as decomposers by digesting food, break down organic matter, alter structurally and chemically, and release it partially in form of waste products.
Temperature & moisture greatly influence microbial activity. Low temperatures and humidity reduce or inhibit microbial activity.
Alternate wetting and drying and continuous dry spells tend to reduce microflora activity and populations
Decomposition rates are highest in warm, wet climates.
The decline in decomposition rate from north to south is a direct result of changes in climate primarily temperature
Diurnal temperature changes effect release of Co2 by respiring decomposer organisms
Absence of fungi (they require oxygen) hinders the decomposition of lignin compounds especially among plants
In mud and sediments of aquatic habitats where oxygen levels can be extremely low, anaerobic carry out most of the decomposition
Diagram illustrating the exchanges between litterbag and soil in a standard litterbag experiment (litterbag- approach use to study dead plant tissues)
Decomposer organisms (bacteria and fungi) colonize the plant litter.
As litter is consumed, a significant proportion of carbon is respired, and nutrients bound in organic matter are mineralized and released to the soil.
To convert plant carbon to microbial biomass, mineral nutrients are taken up in the process of immobilization.
The difference in the rates of mineral nutrient release (mineralization) and immobilization is the rate of net mineralization.
As time progresses, the residual organic matter in the litterbag is composed of a growing proportion of microbial biomass as the original plant material is consumed, respired and converted into microbial biomass.
Immobilize- keeping it from there in an available form, sort of a mineral reserve
During decomposition, not all nutrients will be mineralized in simple inorganic ion form
“whatever will be immobilized today will at some point in the future will be mineralized”
Dead leaf material’s nitrogen content 0.5% to 1.5%
Nitrogen exists in many forms in the soil and is essential for plant growth and development. Most of the nitrogen found in the soil is in numerous different organic forms. However, plants are unable to use organic forms of nitrogen. Normal, rather complex processes in the soil convert nitrogen from organic forms into ammonium.
How can it go above 100%, bacteria/fungi have nitrogen in their own cells, plus they take up what they are breaking down; nitrogen content much higher in decomposers than in plant material
in aquatic systems, the photic zone may be separated from the benthos (where most decomposition goes on)
thus, in aquatic ecosystems, the mineralization of nutrients may not get to the primary producers immediately
Differences in nutrient cycling between terrestrial and open-water aquatic ecosystems
The two zones not linked in deeper waters: physical separation is a major controlling factor controlling the productivity of open-water ecosystems
With the breakdown of the thermocline and mixing of the water column (turnover), nutrients are brought up from the bottom to the surface water. With the onset of spring, increasing temperatures and light in the epilimnion give rise to a peak in productivity due to the increased availability of nutrients in the surface waters. As the spring and summer progress, the nutrients in the surface water are used, reducing the nutrient content of the water and subsequent decline in productivity occurs.
Flowing water has spatial component
Nutrient spiraling
Jack Webster (1975) was the first to point out that because nutrients in streams are subject to downstream transport, there is little nutrient cycling in one place. Water currents move nutrients downstream. Webster suggested that rather than a stationary cycle, stream nutrient dynamics are better represented by a spiral. He coined the term nutrient spiraling to describe stream nutrient dynamics
As an atom of a nutrient completes a cycle within a stream, it may pass through several ecosystem components such as an algal cell, an invertebrate, a fish, or a detrital fragment. Each of these ecosystem components may be displaced downstream by current and therefore contribute to nutrient spiraling. The length of stream required for an atom of a nutrient to complete a cycle is called the spiraling length
Tight spiral- shorter distance of spiral ex. dead leaves & other debris being physically held in place long enough to allow organisms to process the organic matter
Open spiral- longer distance ex. Organisms that shred and fragment organic matter can open the spiral by facilitating the transport of organic materials downstream
Upwelling occurs in the open ocean and along coastlines. The reverse process, called “downwelling,” also occurs when wind causes surface water to build up along a coastline and the surface water eventually sinks toward the bottom.
Water that rises to the surface as a result of upwelling is typically colder and is rich in nutrients. These nutrients “fertilize” surface waters, meaning that these surface waters often have high biological productivity. Therefore, good fishing grounds typically are found where upwelling is common.
In a gas cycle elements move through the atmosphere. Main reservoirs are the atmosphere and the ocean.• In a sedimentary cycle elements move from land to water to sediment. Main reservoirs are the soil and sedimentary rocks.
Leaching occurs when water flowing vertically through the soil transports nutrients in solution downward in the soil profile
Stream-water runoff is often greatest after fire because of reduced water demand for transpiration. High nutrient availability in the soil, coupled with high runoff, can lead to large nutrient losses from the ecosystem
At daylight when photosynthesis begins, plant starts to withdraw CO2 from the air, by afternoon when temperature and relative humidity decreases, rate of photosynthesis declines, CO2 in the atmosphere increases & by sunset, photosynthesis stops, respiration increases therefore atmospheric level of CO2 sharply increases. Same thing happens in aquatic ecosystems
With the onset of the growing season when the landscape is greening, the atmospheric concentration begins to drop as plants withdraw CO2 through photosynthesis. As growing season reaches its end, photosynthesis declines, respiration becomes the dominant process.
For molecular nitrogen (N2) to be used by organisms it must be fixed in a process called fixation this is done by high energy fixation in which factors such as lightening provide the high energy needed to combine nitrogen with the oxygen and hydrogen of water, the second method of fixation is biological fixation which is accomplished by different kinds of micro organisms such as bacteria. Nitrogen is also freed up for use by other organisms by the breakdown of dead organic material in a process called mineralization. In terrestrial ecosystems nitrogen is taken up (assimilation) by plants in the form of ammonia and nitrates, the plants then convert them into amino acids. Herbivores then eat these plants, which convert the obtained nitrogen into different types of amino acids.
The sulfur cycle has both sedimentary and gaseous phases. In the long-term sedimentary phase, sulfur is tied up in organic and inorganic deposits released by weathering and decomposition, and carried to terrestrial ecosystems in salt solution. The gaseous phase of the cycle permits sulfur circulation on a global scale.
Combustion of fossil fuels like gasoline
Volcanic eruption of Mt. Pinatubo where it greatly increased the sulfuric level of the atmosphere
Oceans are a large source of aerosols that contain sulfate but most are redeposited in the ocean as precipitation and dryfall- Dimethylsulfide is the major gas emitted from the oceans
Ozone (O3)- an ambivalent atmospheric gas
At different atmospheric levels:
Stratosphere- shields the planet from UV light; cycling reaction requiring sunlight maintains ozone in this level
Troposphere- born from the union of nitrogen oxides w/ oxygen in the presence of sunlight
Ground level- a damaging pollutant, cutting visibility, irritating eyes and respiratory systems, injuring or killing plant life