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400mya Angiosperms and plant anatomy Million years of evolution:  Cooksonia Practical  # 3 Mid-term evaluations (lab 6 of 10) Lecture Scavenger Hunt Turn in cladistics worksheet p 123-4 Next week: Last lab practical Angiosperm
http://www.johnkyrk.com/evolution.html
Paleo plant and insect interactions
Insects and plants evolved together Older Younger
time Rapid origin
The unique character of angiosperms is that the ovules are completely enclosed in a carpel 1. gametophytes only a few cells 2. immotile sperm- carried to ovule by pollen tube 3. No “spores” because reduced gametophyte
Extinct http://www.pbs.org/wgbh/nova/flower/anat-flash.html Controversy about its floral morphology interpretation  http://www.pbs.org/wgbh/nova/flower/
Knowing the flower anatomy…let’s review the fossil record of earliest angiosperms keep in mind: ,[object Object]
 No consensus about the ancestral relative
 Molecular evidence suggest a Jurassic origin (>150 Ma), but the oldest fossils is early Cretaceous (~125 Ma)MORE INFO: http://www.mobot.org/mobot/research/apweb/welcome.html
UF grad student found this one!
Extinct
Many early flowers are related to living  Angiosperms families
Pollination of ‘primitive’ flowers
Flowers adapted for pollination by "smart" insects
Insects can see uv light
Insects can see uv light
What pollinates these?
Wind-pollinated flowerssecondarily derived in Angiosperms -- flowers are green, small, and often lack petals.   Wind pollinated flowers of deciduous trees species open in early spring – why?
Pick the Pollinator http://www.pbs.org/wgbh/nova/flower/pollinator.html
Plant parts Take a look…
Primitive vs Derived characters http://botany.csdl.tamu.edu/FLORA/tfplab/primder.htm
Fruit and seed dispersal
Don’t forget… Also: ichthyochory
?
What disperses these?
Pleistocene extinction Gomphothere Late Pleistocene Extinctions – 13,000 years ago, N & S America
“Re-wilding of N. America?”
Angiosperms often used for medicine – why? Preparing ayahuasca
3 clades A. Monocotyledones (monocots)  B. Magnoliids C. Eudicotyledones (dicots)
ANGIOSPERMS
Generalized distinguishing characteristics:
A comparison of monocots and dicots  -- know this for lab practical Lab exercise – can you tell them apart?
MONOCOTS All flesh is grass” -- Isaiah Grasses evolved directly with mammals
Bamboo
Banana Pineapple Onion
Palms! ‘ivory palm’ Coconuts Harvesting palm hearts
Bactris gasipaes‘peach palm’
One apical bud!!
phloem xylem Monocot stem -- vascular bundles
Dicot stem Xylem and phloem in rings Dicots have secondary growth
Which tree has a better chance to live? Dicot Monocot (palm)
Dicot stems Illegal mahagony logging Annual growth rings?
We eat many, many dicot fruits Examples? Theobroma cacao
In common? Kale Califlower Kohlrabi Chinese kale cabbage Broccoli Collard greens Brussel sprouts
Brassica oleracea Kale Califlower Kohlrabi Chinese kale cabbage Broccoli Collard greens Brussel sprouts ‘wild mustard’
Brassica oleracea ‘wild mustard’
Vavilov Centersof origin for crop plants http://www.hort.purdue.edu/newcrop/history/lecture05/lec05.html
Anatomy study slides
The life cycle of an angiosperm
Angiosperm life cycle
Plant morphology/anatomy Primary vs. secondary growth A.	Apical meristems - primary growth B.	Cambium (vascular or cork) - secondary growth C.	In lab -. Apical meristems 1. Coleus - stem tip (no. 3) - herbaceous dicot 2. Zea - root longitudinal section (no. 4) - monocot
Seed anatomy Scan
Stems A. Anatomy 1.	parenchyma (pith)		6. xylem 2.	sclerenchyma		7. phloem 3.	epidermis			8. vascular cambium 4.	cork & cork cambium	9. meristem 5.	cortex			10. node B. In lab: 1.	Helianthus - stem (no. 5) - herbaceous dicot 2.	Tilia - stem cross sections (no. 6) - woody dicot 3.	Zea - stem cross section (no. 7)
Morphology of a flowering plant
Morphology of a winter twig
Anatomy of a tree trunk
Organization of primary tissues in young stems
Leaves A. Anatomy 1.	palisade mesophyll		5. stomata 2.	spongy mesophyll		6. xylem 3.	epidermis				7. phloem 4.	cuticle B. In lab: 1.	Ligustrum - leaf section (no. 8)
Leaf anatomy
Simple versus compound leaves a. Simple b. Compound i. palmate ii. pinnate
Roots A. Anatomy 1.	epidermis		5. casparian strip 2.	cortex		           6. pericycle 3.	stele			7. xylem 4.	endodermis		8. phloem B. In lab: 1.	Ranunculus - root (no. 9) 2.	Salix - branch root (no. 10) 3.	radish root hairs 4.	sweet potato demo - storage root 5.	carrot - root stores sugars
Primary growth of a root
Organization of primary tissues in young roots
The formation of lateral roots
Root hairs of a radish seedling
Storage organs A. white potato demo - modified stem B. green onion demo - leaves modified for storage C. celery - leaf petiole modified D. sweet potato demo - storage root E. carrot - root stores sugars
Principal Biological Concepts this lab: A. Angiosperms as dominant plant taxon. B. Ovules enclosed within two integuments and a carpel wall. C. Structure and function of flowers, and importance of pollinators. D. Carpel wall may ripen as pericarp = fruit. E. Monocots versus dicots F. Double fertilization. G. Plant tissues. H. Primary and secondary growth. I. Structure and function of shoots, roots, and leaves.

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