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Photorespiratio
      n
What is
           PHOTORESPIRATION?

• The aberrant use of oxygen by chloroplasts
• An interference with carboxylation caused by
the deviant interaction of RuBisCO with oxygen
•A process that leads to only one 3PGA being
produced in the dark reaction in chloroplasts
*RuBisCO- ribulose 1,5-biphosphate carboxylase/oxygenase
*3PGA- 3-phosphoglyceric acid
Factors that Increases
         Photorespiration




                                      Low CO2
                      High O2      concentration
High temperature   concentration
How
Photorespiration
   Happens??
CH2OP              CH2OP                  CH2OP                    CH2OP

        C=O                 C-OH            HO- C-O-OH             HO- C-O-OH
                                      O2
        C-OH                C-OH                   C=O     H2O     HO- C-OH
        C-OH                C-OH                   C-OH                     C-OH
        CH2OP               CH2OP                  CH2OP                 CH2OP
RuBP
Ribulose-1,5-bisphosphate
             Phospho-         CH2OP
             glycolate
                              COO                                 COO

                                     PO4              3PGA       C-OH

                             CH2OH                               CH2OP
  Peroxisomes                          Glycolate
                             COO
                                                             Calvin Cycle
Glyoxylate
From
Chloroplasts     CH2OH         O2      CHO

                COO                   COO
                                                Mitochondria
               Glycolate       NH3+
                                                  2   CH2NH3
Glycerate        CH2OH              CH2NH3
                                                      COO
                 CHOH               COO

                 COO                Glycine                      CO2
  ATP
                        NAD+                     NH3+
                        NADH
                CH2OH            CH2OH                     CH2OH

                C=O             CHNH3                      CHNH3
  3PGA
                COO             COO                        COO
                                                        Serine
 Hydroxy              Peroxisomes
  pyruvate
From                                                  Serine
                  Phosphoglycolate
Chloroplast




                                          Mitochondria


                             Peroxisome


          Glycolic acid                                  Glycine
Functions of Photorespiration
• Photorespiration Removes Toxic Metabolic
  Intermediates
• Photorespiration Protects from
  Photoinhibition
• Photorespiration Supports Plant Defense
  Reactions
• Photorespiration is Intimately Integrated Into
  Primary Metabolism
Examples of C3 Type of Plants




 Wheat      Soybean     Oat




 Barley    Sunflower   Alfalfa
The End of
  Photorespiration


Next report:
C4 CARBON CYCLE

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Photorespiration

  • 2. What is PHOTORESPIRATION? • The aberrant use of oxygen by chloroplasts • An interference with carboxylation caused by the deviant interaction of RuBisCO with oxygen •A process that leads to only one 3PGA being produced in the dark reaction in chloroplasts *RuBisCO- ribulose 1,5-biphosphate carboxylase/oxygenase *3PGA- 3-phosphoglyceric acid
  • 3. Factors that Increases Photorespiration Low CO2 High O2 concentration High temperature concentration
  • 5.
  • 6. CH2OP CH2OP CH2OP CH2OP C=O C-OH HO- C-O-OH HO- C-O-OH O2 C-OH C-OH C=O H2O HO- C-OH C-OH C-OH C-OH C-OH CH2OP CH2OP CH2OP CH2OP RuBP Ribulose-1,5-bisphosphate Phospho- CH2OP glycolate COO COO PO4 3PGA C-OH CH2OH CH2OP Peroxisomes Glycolate COO Calvin Cycle
  • 7. Glyoxylate From Chloroplasts CH2OH O2 CHO COO COO Mitochondria Glycolate NH3+ 2 CH2NH3 Glycerate CH2OH CH2NH3 COO CHOH COO COO Glycine CO2 ATP NAD+ NH3+ NADH CH2OH CH2OH CH2OH C=O CHNH3 CHNH3 3PGA COO COO COO Serine Hydroxy Peroxisomes pyruvate
  • 8. From Serine Phosphoglycolate Chloroplast Mitochondria Peroxisome Glycolic acid Glycine
  • 9. Functions of Photorespiration • Photorespiration Removes Toxic Metabolic Intermediates • Photorespiration Protects from Photoinhibition • Photorespiration Supports Plant Defense Reactions • Photorespiration is Intimately Integrated Into Primary Metabolism
  • 10. Examples of C3 Type of Plants Wheat Soybean Oat Barley Sunflower Alfalfa
  • 11. The End of Photorespiration Next report: C4 CARBON CYCLE