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  1. 1. Vaucheria Occurrence: It is found in temperate regions and comprises of about 45 species. Most of the species occur in fresh waters (streams, springs and water falls) or found in damp soils. A few species, e.g, Vaucheria pilobolides are marine. Plant body: The thallus consists of branched cylindrical filament. The filament is non septate multinucleated structure. Septa are produced only during the production of reproductive structure. Hepteron: The plant is attached to the substratum by colorless branched, short rhizoids called as hepteron. Cell wall: The cell wall is thin and consists of cellulose and pectin. Vacuole: Tubular filament contains a large central vacuole filled with cell sap. Chloroplast: The chloroplast is small discoid shaped structure without pyrenoids. Reserve food: Reserve food material is in the form of oil drops. Reproduction: Vaucheria reproduces by vegetative, asexual and sexual means. Vegetative Reproduction: Vegetative reproduction in Vaucheria takes place by fragmentation. The filaments are broken accidentally into fragments, each of which develops into a new plant. However, this mode of reproduction is not common. Asexual Reproduction: In aquatic species the asexual reproduction takes place by multiflaggelate zoospores whereas in terrestrial species it is by aplanospores. Aplanospores: Aplanospores are non-motile spores that are formed singly inside structures called aplanosporangium. They are formed during unfavourable conditions at the apical side of aerial hyphae. The aplanosporangium are club-shaped and non-flagellated structures that are separated from the rest of the thallus by a transverse septum.
  2. 2. The aplanospores rupture the cell wall of the aplanosporangium and develop into new filaments. Zoospores: Zoospores are flagellated asexual spores that are born singly inside zoosporangium. Zoosporangia are green-coloured and club-shaped structures that are formed at the apical region of the filaments by forming a cross-wall from the rest of the filament. At the time of the liberation of the spore, the protoplast of the zoosporangium contracts, forming a narrow aperture, from which the zoospores are released. The zoospores become flagellated after liberation. The zoospores are yellowish-green, multiflagellate, multinucleate and ovoid structures that have a small central vacuole. They are also referred to as compound zoospores. Sexual Reproduction:
  3. 3. Sexual reproduction in Vaucheria is of oogamous type. The male sex organ is known as antheridium and the female sex organ is known as oogonium. The plant body of Vaucheria can either be homothallic meaning both the male and female sex organs are born close to each other on the same filament or heterothallic such that male and female sex organs are born on different plants.
  4. 4. Antheridium: A mature antheridium is usually cylindrical structure that develops before oogonium development in most species. It develops close to oogonium. The branch bearing the antheridium becomes densely packed with nuclei and few chloroplasts, therefore the antheridium is often colourless. Later on this branch is cut off from rest of the thallus by a transverse septum. Inside the antheridium, the nuclei divide mitotically and give rise to biflagellated and spindle shaped antherozoids. The flagella are dissimilar. one is whiplash and one is tinsel. The antheridium opens at the apex, antherozoids are released and reached the oogonium. Oogonium: The development of the oogonium starts near the antheridium with a bulging in the main filament. A large mass of colourless cytoplasm called wanderplasm has many nuclei and chloroplasts move into this bulge. The bulge increases in size and becomes an oogonium. The oogonium is separated from the main filament by the formation of a transverse septum. At one side of the oogonium a beak called the receptive spot develops. Apical pore is formed at the apex of the beak through which the antherozoids can enter the oogonium. Fertilization: At the time of fertilization the beak of the oogonium ruptures and an aperture is formed in the antheridium. The oogonium releases a small amount of cytoplasm through the ruptured beak and as a result many antherozoids get stuck in the liquid but only one enters the oogonium. The antherozoids lose their flagella upon coming in contact with the ovum and fuse with it to form a diploid zygote. The zygote secretes a thick wall around it to become an oospore. Germination of oospore: The oospore undergoes a period of rest for some months before germination. On germination it produces a new filament directly
  5. 5. Pinnularia: Habitat: It is a fresh water alga. It is found in pond. It is also present on the moist soil.` Structure: It is unicellular. Its cell is elongated and elliptical. Cell wall: The cell wall is made up of pectic substances. Silica is impregnated in it. So the cell became very hard. Cell wall is composed of two halves called valves. These halves overlap like petridish. The margins of the two valves are covered by a connecting band. The two valves with their inner protoplasts are called frustule. The frustule has two views . The surface view is called valve view and band view is called girdle view . The outer larger valve is called epitheca. The smaller inner valve is called hypotheca. The surface of valve has minute pores or pits . It produces characteristic markings on the surface of valves . These markings are bilaterally symmetrical. The marking free area is called axial field . The markings are arranged in linear rows .
  6. 6. The axial field contains a longitudinal slit called raphe. The raphe is not continuous throughout the length of the valve . The raphe is wedge shaped in transverse section. It has circular thickening called central nodules in the centre. Similar thickening also present at the outer ends of raphe . These are called polar nodules . These nodules open to the external aqueous medium
  7. 7. Cytoplasm: Cytoplasm is arranged in a parietal layer. A large central vacuole is present inside the cell. Nucleus: The single nucleus is suspended in the centre of vacuole by a cytoplasmic bridge. Chloroplast: Two chloroplast are present along the two sides of the cells. They contain chlorophyll a, c, beta carotene, fucoxanthin and diatomin pigment. Fucoxanthin gives the alga characteristic colour. The plastids are few in number. Excess food is stored in the form of oil or chrysolaminarin. Locomotion: Locomotory organs are absent in pinnularia. It moves by gliding movements. Circulation of the streaming cytoplasm causes this movement. Mucilage helps in this gliding. Reproduction: Vegetative and sexual reproduction are present in it .
  8. 8. Vegetative reproduction: It is the most common method of reproduction. It produces daughter cells slightly different in sizes. Vegetative reproduction occurs by simple cell division. The protoplast expands. It causes slight separation of epitheca and hypotheca. Nuclear division occurs by mitosis and cell divides into two parts. Each half receives one half of the parent cell. It synthesizes new valve. New valve is fit into the parent valve. Thus new valves are always smaller than the parent halves. Thus one generation gradually become smaller in size. It reacted to minimum size. Then its size is restored by auxospore formation. But second generation remains of same size. Sexual reproduction (Auxospore formation).
  9. 9. 1. Gamete formation: In some species of Pinnularia, two cells from common parent or different parents envelope in a common mucilaginous sheath. The nuclei of both cells divide by meiosis to form four nuclei. Three nuclei disintegrate. The fourth one enlarges. Its protoplast metamorphosed into gamete. The gametes are liberated from the parent frustules. They fuse to form zygote. The zygote enlarges to form auxospore. The auxospore secretes new valve and become adult. 2. Parthenogenesis: The protoplast of cell secretes a large amount of mucilage. Therefore two valves are separated from each other. It separates the inner protoplast. The protoplast grows to its maximum size. It secretes silica rich pectic membrane. New valves are secreted inside the membrane. Nuclear division does not occur. Therefore auxospores are formed by parthenogenesis.
  10. 10. Ectocarpus: Occurrence: It is a marine brown alga. It is abundantly found in cold water. A few species occur in fresh water. The plant grows attached to rocks and stones along coasts. Some species are epiphytes on other algae like members of Fucale’s and laminaria. Plant body: Plant body is filamentous and branched. There are two types of filaments, the prostrate and erect filament. Prostrate filament: The prostrate system is branched and attached to the substratum with the help of rhizoids. In epiphytic species this system occasionally penetrates the host. In free floating species the prostrate system is poorly developed. Erect filament: The erect branches are arranged end-to-end, such that they are polysiphonous in nature. Sometimes, the erect branches may grow vertically to form more than one row of cells.
  11. 11. Growth of thallus: The growth of thallus in primitive brown algae is by an intercalary meristem at the base of a hair or filament. This type of growth is called trichothallic growth. Cell structure: The cells are rectangular or cylindrical. Cell wall is differentiated into outer pectic and inner cellulosic layers. The outer layer becomes slimy or gummy due to presence of pectic compounds. The most important constituent of the cell wall is alginic acid found in the outer layer of the cell wall. It forms a substance algin which is important economically. The reserve food is mannitol and laminarin starch. It is golden brown in colour due to presence of dominant fucoxanthin. The other photosynthetic pigments are chlorophyll a, c, beta carotene and other xanthophylls. The protoplast contains one central nucleus and many chromatophores. The number and shape of chromatophore varies with species. Pyrenoid are associated with chromatophores.
  12. 12. Reproduction: Ectocarpus reproduces asexually by zoospores and sexually by isogamy or anisogamy. The oogamous type of sexual reproduction is absent. Asexual Reproduction: The asexual reproduction takes place by zoospores. The zoospores are biflagellate. The diploid plant (2n) develops two types of sporangia. unilocular sporangia and plurilocular sporangia. The unilocular sporangia develop haploid zoospores and plurilocular sporangia develop diploid zoospores. Unilocular Sporangia:The unilocular sporangia develop from the apical cell of short lateral branches. The cell enlarges and functions as sporangial initial. The nucleus of the sporangium undergoes meiosis followed by several mitotic divisions to produce 32-64 daughter nuclei. These nuclei accumulate some cytoplasm and develop individual units. Each unit metamorphoses into a uninucleate and biflagellate zoospore also called meio
  13. 13. zoospore. The flagella are unequal and laterally inserted. During liberation the apex of the sporangium wall gets dissolved and the haploid zoospores are liberated in a gelatinous mass. After some time they get free from the gelatinous mass and swim freely in water. After the liberation of zoospores a new sporangium may be produced within the old sporangial wall. Plurilocular Sporangia: These are elongated multicellular structures that develop at the tip of short lateral branches. The sporangial initial becomes enlarged and undergoes repeated mitotic divisions to produce a vertical row of 6-12 cells. The cells then undergo several vertical divisions thus a multi chambered structure is formed i.e., the plurilocular sporangium. The protoplast of each cell metamorphoses into diploid biflagellated zoospore. The zoospores are liberated through a terminal or a lateral pore formed in the wall of the sporangium. Sexual Reproduction: Majority of the species are isogamous and homothallic. The gametes are produced inside the plurilocular gametangia, developed on haploid plants.
  14. 14. Plurilocular Gametangia: These are large, elongated and multicellular structures. Both plurilocular gametangia and plurilocular sporangia are alike. The plurilocular gametangia produce haploid gametes while plurilocular sporangia produce diploid zoospores. The gametes are slightly smaller in size than the zoospores. The gametes are liberated from the gametangia following the same procedure as that of zoospore liberation from the plurilocular sporangia. Fertilization and Germination of zygote: Fertilization results in a diploid zygote. The zygote germinates into a diploid sporophyte without undergoing any resting period. The sporophytic plant again develops unilocular and plurilocular sporangi

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