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AGB 22012: Crop pests and Management
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Insect pest of Rice
a) Stem borers
1. Yellow stem borer, Scirpophaga incertula
2. Gall fly or Gall midge, Orseolia oryzae
GM
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Insect pest of Rice
b) Leaf feeders
1.Whorl maggot, Hydrellia sasakii
2. Swarming caterpillar, Spodoptera mauritia
3. Case worm, Nymphula depunctalis
4. Leaf folder, Cnaphalocrosis medinalis
Marasmia patnalis
5. Skipper butterfly, Pelopides mathias
GM
4
Insect pest of Rice
b) Leaf feeders …
6. Green horned caterpillar,
Melanitis leda ismene
7. Hispa or spiny beetle, Dicladispa armigera
8. Grass hopper,
Oxya nitidula
Hieroglyphus banyan
GM
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Insect pest of Rice
c) Sap feeders
1. Thrips, Stencheatothrips biformis
2. Green leafhopper, Nephotettix nigropictus
N. cincticeps , N. impicticeps
3. White leaf hopper, Cofana spectra
4. Zig zag leaf hopper, Recilia dorsalis
5. Brown plant hopper, Nilaparvata lugens
6. Ear head bug, Leptocorisa acuta
GM
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Yellow stem borer
Scirpophaga incertula
Lepidoptera : Pyralidae
White ear Dead heart
GM
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Yellow stem borer
Scirpophaga incertulas
Lepidoptera : Pyralidae
White ear Dead heart
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Scirpophaga incertula
Excreta indicate
the holes
Stem hole with larva Stem with larva and excreta
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Scirpophaga incertula
Female
Male
Egg
Larva
pupa
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Fecundity:
•15-80 eggs/Adult
- in mass near tip on upper surface of
tender leaf blade during night and cover
them with buff coloured hairs and scales –
derived from anal tuft.
• 2-3 egg masses /female
• Incubation period : 5 – 8 days
GM
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• One larva bores into the stem near nodal region
• Usually 1 larva/stem, but 2-4 larvae/stem also
reported
• Larva – Migratory
• come out of stem and prepare an enclosing case
• infest new plant
GM
• Pale white with dark brown head and
prothoracic shield
• Enter leaf sheath and feed for 2-3 days
Larva:
12
Larva.. Cont..
• Larval period – 33 - 41 days
• length – 20 mm
• Before pupation, larva plug the exit hole
GM
Pupa: in silken cocoon
• Dark brown
• 12 mm long
• Pupal period - 6-10 days (may get prolonged
depending upon the weather condition)
13
Moth
• Attracted to light
• Female:male – 2:1
• Cold weather
• high humidity &
• low temperature
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Natural enemies
Egg Parasitoids
• Tetrastichus sp.
• Telenomus beneficians
• Trichogramma sp.
Larval parasites
Amauromorpha accepta
Isotima javensis
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Hollow cavity in the rice stem
Symptoms of Rice gall midge
Orseolia oryzae
Diptera : Cecidomyiidae
Silver shoots of gall midge larvae.
GM
Onion Leaf
16
Adult
eggs
Maggot
17
pupa
Rice Whorl Maggot
Hydrellia sesakii
Diptera : Ephyridae
egg
Adult
Maggot
GM
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Symptoms of Rice Whorl Maggot
Hydrellia sesakii
Diptera : Ephyridae
Yellow spots and streaks on leaves
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egg larva
Rice case worm
Nymphula depunctalis
Lepidoptera : Pyralidae
adult
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AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
20
Damage field leaves dry up and turn white
Floating larval case
Leaf blades are cut as by scissors
Larval head come out from case
Rice case worm
Nymphula depunctalis
Lepidoptera : Pyralidae
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AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
21
adult
Spineless adult
pupa
grub
Rice Hispa beetle
Dicladispa armigera
Coleoptera : Chrysomelidae
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
22
Damage of Rice Hispa beetle
Dicladispa armigera
Coleoptera : Chrysomelidae
Leaf mining
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Swarming caterpillar/ Army worm
Spodoptera mauritia
Lepidoptera : Noctuidae
Adults feeding on the
plants
Remove the leaf blades
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
24
Swarming caterpillar/ Army worm
Spodoptera mauritia
Lepidoptera : Noctuidae
Egg masses
covered with hairs
Larva hatching from
the egg masses
Larva
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
25
Pupate on the plant/ into the soil
Adult
Swarming caterpillar/ Army worm
Spodoptera mauritia
Lepidoptera : Noctuidae
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Ear cutting caterpillar (Army worm)
Mythimna separata
Lepidoptera : Noctuidae
Plant damage
Egg mass
Egg mass folded in leaf
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Ear cutting caterpillar (Army worm)
Mythimna separata
Lepidoptera : Noctuidae
Larva
pupa
Adult
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Cut worm
Spodoptera litura
Lepidoptera : Noctuidae
Larva
Adult
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AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Damage
Leaf folder
Lepidoptera : Pyralidae
Folded leaves
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AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
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Leaf folder
Cnaphalocrocis medinalis
Lepidoptera : Pyralidae
Eggs
Larva
pupa
adult
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
31
Leaf folder
Marasmia patnalis
Lepidoptera : Pyralidae
Adult
Larva
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
32
Thrips
Stenchaetothrips biformis
Thysanoptera : Thripidae
Damage
Larva
Mature larva
Adult
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
33
Green horned caterpillar
Melanitis leda ismene
Lepidoptera : Satyridae
Adult
Egg
Larva
pupa Damage
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
34
Green horned caterpillar
Mycalesis sp.
Lepidoptera : Satyridae
Adult
Larvae
pupa
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
35
Skipper butterfly
Pelopidas mathias
Lepidoptera : Hesperiidae
adult
egg
larva
pupa
Leaf tip folded
symptom
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
36
Hairy caterpillar
Creatonotus gangis
Lepidoptera : Arctiidae
Larva
pupa
Adult
Egg mass
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
37
Oxya
Acrida
Atractomorpha
Rice Grasshopper
Orthoptera : Acrididae
Egg covered with frothy secretions
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
38
Green leaf hoppers
Nephotettix virescens
Hemiptera : Cicadellidae
Nymph Adult
eggs
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
39
Green leaf hoppers
Nephotettix nigropictus
Hemiptera : Cicadellidae
Nymph Adult
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
40
Adult Nymph
Nephotettix malayanus
Nephotettix cincticeps
Adult Nymph
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
41
Zig zag hoppers
Recilia dorsalis
Hemiptera : Cicadellidae
Adult
Nymph
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
42
White leaf hopper
Cofana spectra
Hemiptera : Cicadellidae
Egg
Adult
Nymph
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
43
Brown Plant hopper
Nilaparvata lugens
Hemiptera : Delphacidae
Adult
Egg Nymph
Hopper burn Hopper burn
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
44
White backed Plant hopper
Sogatella furcifera
Hemiptera : Delphacidae
Male
Female
Egg
Nymph
Hopper burn
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
45
Rice ear head bug
Leptocorisa acuta
Hemiptera : Alydidae
Adult Nymph
egg
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
46
Damage grain
Rice ear head bug
Leptocorisa acuta
Hemiptera : Coreidae
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
47
Sting bug
Nezara viridula
Hemiptera : Pentatomidae
eggs Nymph
Adult
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
48
Sting bug
Dolicorus indicus
Hemiptera : Pentatomidae
Adult
Eggs & Nymphs
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
49
Root aphid
Rhopalosiphum rufiabdominalis
Hemiptera : Aphididae
Yellowing
adult
Nymphs
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
50
Mole cricket
Gryllotalpa orientalis
Orthoptera : Gryllotalpidae
Damage
Adult
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
51
White grub
Holotrichia sp.
Coleoptera :Scarabaeidae
Damage
Grub
pupa
adult
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
52
Black beetle
Phyllophaga sp.
Coleoptera :Scarabaeidae
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
53
Termite
Macrotermes gilvus
Isoptera : Termitidae
Adult
Damage
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
54
Ants
Solenopsis geminata
Hymenoptera : Formicidae
Adult
Reduce seed
germination
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
55
End of Part 1:Insect pests
GM
AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
Acknowledgement:
1. International Rice Research Institute, Philippines
2. Department of Agriculture, Ministry of Agriculture, Sri Lanka

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Rice insect pests - GM

  • 1. 1 AGB 22012: Crop pests and Management
  • 2. 2 Insect pest of Rice a) Stem borers 1. Yellow stem borer, Scirpophaga incertula 2. Gall fly or Gall midge, Orseolia oryzae GM
  • 3. 3 Insect pest of Rice b) Leaf feeders 1.Whorl maggot, Hydrellia sasakii 2. Swarming caterpillar, Spodoptera mauritia 3. Case worm, Nymphula depunctalis 4. Leaf folder, Cnaphalocrosis medinalis Marasmia patnalis 5. Skipper butterfly, Pelopides mathias GM
  • 4. 4 Insect pest of Rice b) Leaf feeders … 6. Green horned caterpillar, Melanitis leda ismene 7. Hispa or spiny beetle, Dicladispa armigera 8. Grass hopper, Oxya nitidula Hieroglyphus banyan GM
  • 5. 5 Insect pest of Rice c) Sap feeders 1. Thrips, Stencheatothrips biformis 2. Green leafhopper, Nephotettix nigropictus N. cincticeps , N. impicticeps 3. White leaf hopper, Cofana spectra 4. Zig zag leaf hopper, Recilia dorsalis 5. Brown plant hopper, Nilaparvata lugens 6. Ear head bug, Leptocorisa acuta GM
  • 6. 6 Yellow stem borer Scirpophaga incertula Lepidoptera : Pyralidae White ear Dead heart GM
  • 7. 7 Yellow stem borer Scirpophaga incertulas Lepidoptera : Pyralidae White ear Dead heart GM
  • 8. 8 Scirpophaga incertula Excreta indicate the holes Stem hole with larva Stem with larva and excreta GM
  • 10. 10 Fecundity: •15-80 eggs/Adult - in mass near tip on upper surface of tender leaf blade during night and cover them with buff coloured hairs and scales – derived from anal tuft. • 2-3 egg masses /female • Incubation period : 5 – 8 days GM
  • 11. 11 • One larva bores into the stem near nodal region • Usually 1 larva/stem, but 2-4 larvae/stem also reported • Larva – Migratory • come out of stem and prepare an enclosing case • infest new plant GM • Pale white with dark brown head and prothoracic shield • Enter leaf sheath and feed for 2-3 days Larva:
  • 12. 12 Larva.. Cont.. • Larval period – 33 - 41 days • length – 20 mm • Before pupation, larva plug the exit hole GM Pupa: in silken cocoon • Dark brown • 12 mm long • Pupal period - 6-10 days (may get prolonged depending upon the weather condition)
  • 13. 13 Moth • Attracted to light • Female:male – 2:1 • Cold weather • high humidity & • low temperature
  • 14. 14 Natural enemies Egg Parasitoids • Tetrastichus sp. • Telenomus beneficians • Trichogramma sp. Larval parasites Amauromorpha accepta Isotima javensis
  • 15. 15 Hollow cavity in the rice stem Symptoms of Rice gall midge Orseolia oryzae Diptera : Cecidomyiidae Silver shoots of gall midge larvae. GM Onion Leaf
  • 17. 17 pupa Rice Whorl Maggot Hydrellia sesakii Diptera : Ephyridae egg Adult Maggot GM
  • 18. 18 Symptoms of Rice Whorl Maggot Hydrellia sesakii Diptera : Ephyridae Yellow spots and streaks on leaves GM
  • 19. 19 egg larva Rice case worm Nymphula depunctalis Lepidoptera : Pyralidae adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 20. 20 Damage field leaves dry up and turn white Floating larval case Leaf blades are cut as by scissors Larval head come out from case Rice case worm Nymphula depunctalis Lepidoptera : Pyralidae GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 21. 21 adult Spineless adult pupa grub Rice Hispa beetle Dicladispa armigera Coleoptera : Chrysomelidae GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 22. 22 Damage of Rice Hispa beetle Dicladispa armigera Coleoptera : Chrysomelidae Leaf mining GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 23. 23 Swarming caterpillar/ Army worm Spodoptera mauritia Lepidoptera : Noctuidae Adults feeding on the plants Remove the leaf blades GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 24. 24 Swarming caterpillar/ Army worm Spodoptera mauritia Lepidoptera : Noctuidae Egg masses covered with hairs Larva hatching from the egg masses Larva GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 25. 25 Pupate on the plant/ into the soil Adult Swarming caterpillar/ Army worm Spodoptera mauritia Lepidoptera : Noctuidae GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 26. 26 Ear cutting caterpillar (Army worm) Mythimna separata Lepidoptera : Noctuidae Plant damage Egg mass Egg mass folded in leaf GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 27. 27 Ear cutting caterpillar (Army worm) Mythimna separata Lepidoptera : Noctuidae Larva pupa Adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 28. 28 Cut worm Spodoptera litura Lepidoptera : Noctuidae Larva Adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 29. 29 Damage Leaf folder Lepidoptera : Pyralidae Folded leaves GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 30. 30 Leaf folder Cnaphalocrocis medinalis Lepidoptera : Pyralidae Eggs Larva pupa adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 31. 31 Leaf folder Marasmia patnalis Lepidoptera : Pyralidae Adult Larva GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 32. 32 Thrips Stenchaetothrips biformis Thysanoptera : Thripidae Damage Larva Mature larva Adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 33. 33 Green horned caterpillar Melanitis leda ismene Lepidoptera : Satyridae Adult Egg Larva pupa Damage GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 34. 34 Green horned caterpillar Mycalesis sp. Lepidoptera : Satyridae Adult Larvae pupa GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 35. 35 Skipper butterfly Pelopidas mathias Lepidoptera : Hesperiidae adult egg larva pupa Leaf tip folded symptom GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 36. 36 Hairy caterpillar Creatonotus gangis Lepidoptera : Arctiidae Larva pupa Adult Egg mass GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 37. 37 Oxya Acrida Atractomorpha Rice Grasshopper Orthoptera : Acrididae Egg covered with frothy secretions GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 38. 38 Green leaf hoppers Nephotettix virescens Hemiptera : Cicadellidae Nymph Adult eggs GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 39. 39 Green leaf hoppers Nephotettix nigropictus Hemiptera : Cicadellidae Nymph Adult AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 40. 40 Adult Nymph Nephotettix malayanus Nephotettix cincticeps Adult Nymph AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 41. 41 Zig zag hoppers Recilia dorsalis Hemiptera : Cicadellidae Adult Nymph AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 42. 42 White leaf hopper Cofana spectra Hemiptera : Cicadellidae Egg Adult Nymph GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 43. 43 Brown Plant hopper Nilaparvata lugens Hemiptera : Delphacidae Adult Egg Nymph Hopper burn Hopper burn GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 44. 44 White backed Plant hopper Sogatella furcifera Hemiptera : Delphacidae Male Female Egg Nymph Hopper burn GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 45. 45 Rice ear head bug Leptocorisa acuta Hemiptera : Alydidae Adult Nymph egg GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 46. 46 Damage grain Rice ear head bug Leptocorisa acuta Hemiptera : Coreidae GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 47. 47 Sting bug Nezara viridula Hemiptera : Pentatomidae eggs Nymph Adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 48. 48 Sting bug Dolicorus indicus Hemiptera : Pentatomidae Adult Eggs & Nymphs GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 49. 49 Root aphid Rhopalosiphum rufiabdominalis Hemiptera : Aphididae Yellowing adult Nymphs GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 50. 50 Mole cricket Gryllotalpa orientalis Orthoptera : Gryllotalpidae Damage Adult GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 51. 51 White grub Holotrichia sp. Coleoptera :Scarabaeidae Damage Grub pupa adult AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 52. 52 Black beetle Phyllophaga sp. Coleoptera :Scarabaeidae AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 53. 53 Termite Macrotermes gilvus Isoptera : Termitidae Adult Damage AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 54. 54 Ants Solenopsis geminata Hymenoptera : Formicidae Adult Reduce seed germination GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan
  • 55. 55 End of Part 1:Insect pests GM AGB 3201 :Plant Protection and Epidemiology – Rice ecosystem-Insect pests Dr.G.Mikunthan Acknowledgement: 1. International Rice Research Institute, Philippines 2. Department of Agriculture, Ministry of Agriculture, Sri Lanka