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Biology and Physiology
          of
Pomacea Canaliculata



          Presented by
CHANG PEI XIAN       D 20101037455
Pomacea     Canaliculata are amphibious, both
 in physiology and behaviour.
Their mantle cavity contains both a ctenidium
 (gill) and a portion modified as a pulmonary
 sacs or “lung”.
have gills, but can survive for extended
 periods exposed to air.
Have a trap door (operculum) to protect their
 soft tissues when withdrawn into the shell.
Coloration   of the shell is brown or olive
 brown, often with darker bands and blotches.
Show sexual dimorphism in shell size, shape,
 thickness and weight.
They have been reported to change sex. Sex
 change is from male to female (during a period
 of without aestivation)
Shows variation related mainly to different
 thermal regimes.
Pomecea    Canaliculata have long life cycles
 (>3 years but depends on the thermal regimes).
 Iteropareous in regions with hydric and
 thermal seasonality.
When reared at a constant temperature of
 25 C, P. Canaliculata’s life span reduced to
 less than 14 months.
Growth is not continuous but seasonal.
 temperature appears to be the main factor
 affecting both the maturation process and their
 activity.
Sexually mature   when it is 25 mm.
Able to tolerate low levels of salinity.
Experimental traits have shown that growth
 rates, longevity and age at maturity are highly
 dependent on temperature, crowding and food
 availability.
Mobility and feeding are highly reduced when
 water     depth    is    lower      than   shell
 diameter, drought survival is probably related
 to size.
Anatomy of Pomacea Canaliculata
 Male Snail
Female Snail
Digestive System
 The  digestive system of Pomacea Canaliculata is
  adapted to feed on aquatic plants.
 It can be divided in five regions:
 The intake region
   (mouth and buccal cavity with radula and jaws)
 The pre-digestion region
  (esophagus with salivary glands and lateral pouches)
 Digestion region
  (3 chambered stomach with associated digestive glands)
 Uptake region
  (intestine)
 The excretion region
  (rectum and anus)
Scheme of the Digestive Tract
 Mouth
 vertical slit opening, located between the labial
  tentacles and leading to the buccal cavity.
 Food is located with the labial tentacles and when
  needed gathered from the water surface with foot.




 The radula of an eating snail in   The radula knobs and jaws of an eating
 Detail(Pomacea Canaliculata)       Snail(Pomacea Canaliculata)
Buccal   cavity
 A muscular cavity with a set of calcareous
  jaws, 2 radula knobs and the radula (rasp
  tongue), is situated behind the mouth opening.
 When the radula is bend and stretched, this
  spreading the teeth on it, which provides the
  grasp function.
 After the food has pulled into the buccal
  cavity, calcareous jaws cut off the piece of food.
Stomach
 A pink “U” shaped structure on the left side of the
   body whorl.
 Consists of three compartments:
 ◦ the muscular gizzard (posterior chamber)
 ◦ the vestibule (anterior chamber)
 ◦ the style sac.
Intestine
 Lies close to the style sac and coils around the
  posterior kidney.
 Can be divided in several parts: the caecum, the
  first, second and third part of the intestine and anal
  papilia.
 The digestion of the food is partially done by
  adding enzymes by the snail itself and partly by the
  micro-organisms in here.
 The uptake of the food compound is carry out
  through the walls of the intestine and transported to
  the blood stream.
Sensory System
Pomacea    Canaliculata are highly dependent
 on their smell sense.
Able to locate food and recognize other snails
 of their own species with their smell sense.
Vision is weak; functions merely as a light
 direction detector.
The tactile sense is well developed.
The hearing capabilities are worthless, even
 more, they are completely deaf.
Body:
 the whole body surface contains chemo- and
  mechanoreceptors.

Eyes:




     The eye on the eye stalk of a Pomacea Canaliculata Snail.
  The eyes of the Pomacea Canaliculata are located
   at the base of the tentacles, on top of the eyestalks.
The structure of the eye does not provide
 detail vision, they rather function as
 directional light sensors.
Colour vision is absent; Pomacea
 Canaliculata is colour-blind.
Two mechanisms are used in their eyes to
 increase sensitivity to light: the eyes are
 relatively large with a large lens and the light
 sensitive cells (rhabdoms) are grouped and
 connected with the same neuron.
 Osphridia:
 A chemosensory structure that is located in the mantle
  cavity, in front on the lung.
 Gives the snail capability to smell chemical substances
  in the water.
Statocysts:
 Function as balancing organs, used by the
  snail to detect its position with regard to the
  ground.
 Located inside the snail’s body closed to the
  pedal ganglia.

Tentacles:
 Important sensory organs.
 Pomacea Canaliculata highly rely on the
  smell capacity and the sensitivity of their
  tentacles to navigate in their environment.
Nervous System
Central nervous system consists of a several
 ganglia connected with each other by a
 commisura.
The perikaya (cell bodies of the nerve cells)
 are concentrated in these ganglia.
Some parts of the nervous system are
 symmetric while others are not.
There isn’t a central brain in snails like
 mammals have.
Scheme of the Nervous System
Respiratory System
The   respiratory system of the Pomacea
 Canaliculata : a combination of a gill and a
 lung.
The arise of the lung in Pomacea Canaliculata
 can be explained by the following conditions:
 Low oxygen level in the tropical pools.
 Use during periods of drought.
 Increases the action radius of the snail.
Retained   their gill during the course of
 evolution, this is contrary to the Pulmonata
 snails, which only have a lung and no gills.
Having a gill provide a big advantage - can
 avoid attacks by the land predators.
Have a siphon (breathing tube) so that they
 can breath air while staying hidden below the
 surface.
Description of the Respiratory Organs
 Ctenidium:
 Gill of the Pomacea Canaliculata.
 Consists of a row of gill filaments that hang
  into the mantle cavity.

 Osphradius:
 Smell organ of Pomacea Canaliculata.
Lung:
An air filled sac in the left mantle cavity.
Walls of the pulmonary sac are well supplied
 with blood to exchange oxygen and carbon
 dioxide.

Siphon:
An organ consisting of a fold of the mantle
  cavity.
Enable the snail to breath air while staying
  submerged.
Heart and Circulation
There is only one auricle that receives oxygen
 rich blood from the lung and gills,
 deoxygenated blood from kidney.
No separated blood circulation for oxygen rich
 and deoxygenated blood.
Blood has two functions:
 As hydroskeleton(structural support).
 Transport of O2, CO2, hormones, nutrition and
  waste products.
Open    circulation.
H = Head - cephalic hemocoel    Au = Auricle
F = Foot hemocoel               V = Ventricle
E = Oesophagus                  Aa = Aorta anterior
Vm = Visceral mass (hemocoel)   Ap = Aorta posterior
aK = anterior (front) Kidney    Vs = Visceral sinus
pK = posterior (back) Kidney    Fs = Foot sinus
A = Ampulla
The  transport of the blood to and from the
 organs occur through artery.
They don’t have capillary veins and arterioles.
Reproductive System
Has  separated sexes (dioecious).
The differences between the sexes are not
 visible at shell colour or pattern.
Male reproductive system
 The  male reproductive tract consist of the testis
  and vas deferens, the seminal vesicle and the
  prostate gland, the penial sac, the penis and the
  penis sheath.
 During mating activities, this penial papilla bends
  towards the sperm groove in which the sperm is
  conducted.
 The sperm is conveyed in the penial duct at the
  base of the penis.
 On erection the penis comes out of the penial sac
  and is grasped around on the lower part by a
  muscular penial sheath from the mantle.
Female Reproductive System
The   egg cells are stored in the ovaries.
To produce an egg, the egg cell is brought to
 the receptaculum seminis. In this place, the
 fertilization of the egg cells takes place.
The spermatozoa (sperm cells) of the male can
 survive for more than a month in the
 female, and several egg clutches can be
 fertilized with a single copulation.
The eggs were deposited above the water line.
Biology and physiology of Pomacea Canaliculata

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Biology and physiology of Pomacea Canaliculata

  • 1. Biology and Physiology of Pomacea Canaliculata Presented by CHANG PEI XIAN D 20101037455
  • 2. Pomacea Canaliculata are amphibious, both in physiology and behaviour. Their mantle cavity contains both a ctenidium (gill) and a portion modified as a pulmonary sacs or “lung”. have gills, but can survive for extended periods exposed to air. Have a trap door (operculum) to protect their soft tissues when withdrawn into the shell.
  • 3. Coloration of the shell is brown or olive brown, often with darker bands and blotches. Show sexual dimorphism in shell size, shape, thickness and weight. They have been reported to change sex. Sex change is from male to female (during a period of without aestivation) Shows variation related mainly to different thermal regimes.
  • 4. Pomecea Canaliculata have long life cycles (>3 years but depends on the thermal regimes).  Iteropareous in regions with hydric and thermal seasonality. When reared at a constant temperature of 25 C, P. Canaliculata’s life span reduced to less than 14 months. Growth is not continuous but seasonal.  temperature appears to be the main factor affecting both the maturation process and their activity.
  • 5. Sexually mature when it is 25 mm. Able to tolerate low levels of salinity. Experimental traits have shown that growth rates, longevity and age at maturity are highly dependent on temperature, crowding and food availability. Mobility and feeding are highly reduced when water depth is lower than shell diameter, drought survival is probably related to size.
  • 6. Anatomy of Pomacea Canaliculata Male Snail
  • 8. Digestive System  The digestive system of Pomacea Canaliculata is adapted to feed on aquatic plants.  It can be divided in five regions: The intake region (mouth and buccal cavity with radula and jaws) The pre-digestion region (esophagus with salivary glands and lateral pouches) Digestion region (3 chambered stomach with associated digestive glands) Uptake region (intestine) The excretion region (rectum and anus)
  • 9. Scheme of the Digestive Tract
  • 10.  Mouth vertical slit opening, located between the labial tentacles and leading to the buccal cavity. Food is located with the labial tentacles and when needed gathered from the water surface with foot. The radula of an eating snail in The radula knobs and jaws of an eating Detail(Pomacea Canaliculata) Snail(Pomacea Canaliculata)
  • 11. Buccal cavity A muscular cavity with a set of calcareous jaws, 2 radula knobs and the radula (rasp tongue), is situated behind the mouth opening. When the radula is bend and stretched, this spreading the teeth on it, which provides the grasp function. After the food has pulled into the buccal cavity, calcareous jaws cut off the piece of food.
  • 12. Stomach A pink “U” shaped structure on the left side of the body whorl. Consists of three compartments: ◦ the muscular gizzard (posterior chamber) ◦ the vestibule (anterior chamber) ◦ the style sac.
  • 13. Intestine Lies close to the style sac and coils around the posterior kidney. Can be divided in several parts: the caecum, the first, second and third part of the intestine and anal papilia. The digestion of the food is partially done by adding enzymes by the snail itself and partly by the micro-organisms in here. The uptake of the food compound is carry out through the walls of the intestine and transported to the blood stream.
  • 14. Sensory System Pomacea Canaliculata are highly dependent on their smell sense. Able to locate food and recognize other snails of their own species with their smell sense. Vision is weak; functions merely as a light direction detector. The tactile sense is well developed. The hearing capabilities are worthless, even more, they are completely deaf.
  • 15. Body: the whole body surface contains chemo- and mechanoreceptors. Eyes: The eye on the eye stalk of a Pomacea Canaliculata Snail. The eyes of the Pomacea Canaliculata are located at the base of the tentacles, on top of the eyestalks.
  • 16. The structure of the eye does not provide detail vision, they rather function as directional light sensors. Colour vision is absent; Pomacea Canaliculata is colour-blind. Two mechanisms are used in their eyes to increase sensitivity to light: the eyes are relatively large with a large lens and the light sensitive cells (rhabdoms) are grouped and connected with the same neuron.
  • 17.  Osphridia: A chemosensory structure that is located in the mantle cavity, in front on the lung. Gives the snail capability to smell chemical substances in the water.
  • 18. Statocysts: Function as balancing organs, used by the snail to detect its position with regard to the ground. Located inside the snail’s body closed to the pedal ganglia. Tentacles: Important sensory organs. Pomacea Canaliculata highly rely on the smell capacity and the sensitivity of their tentacles to navigate in their environment.
  • 19. Nervous System Central nervous system consists of a several ganglia connected with each other by a commisura. The perikaya (cell bodies of the nerve cells) are concentrated in these ganglia. Some parts of the nervous system are symmetric while others are not. There isn’t a central brain in snails like mammals have.
  • 20. Scheme of the Nervous System
  • 21. Respiratory System The respiratory system of the Pomacea Canaliculata : a combination of a gill and a lung. The arise of the lung in Pomacea Canaliculata can be explained by the following conditions: Low oxygen level in the tropical pools. Use during periods of drought. Increases the action radius of the snail.
  • 22. Retained their gill during the course of evolution, this is contrary to the Pulmonata snails, which only have a lung and no gills. Having a gill provide a big advantage - can avoid attacks by the land predators. Have a siphon (breathing tube) so that they can breath air while staying hidden below the surface.
  • 23. Description of the Respiratory Organs Ctenidium: Gill of the Pomacea Canaliculata. Consists of a row of gill filaments that hang into the mantle cavity. Osphradius: Smell organ of Pomacea Canaliculata.
  • 24. Lung: An air filled sac in the left mantle cavity. Walls of the pulmonary sac are well supplied with blood to exchange oxygen and carbon dioxide. Siphon: An organ consisting of a fold of the mantle cavity. Enable the snail to breath air while staying submerged.
  • 25. Heart and Circulation There is only one auricle that receives oxygen rich blood from the lung and gills, deoxygenated blood from kidney. No separated blood circulation for oxygen rich and deoxygenated blood. Blood has two functions: As hydroskeleton(structural support). Transport of O2, CO2, hormones, nutrition and waste products. Open circulation.
  • 26. H = Head - cephalic hemocoel Au = Auricle F = Foot hemocoel V = Ventricle E = Oesophagus Aa = Aorta anterior Vm = Visceral mass (hemocoel) Ap = Aorta posterior aK = anterior (front) Kidney Vs = Visceral sinus pK = posterior (back) Kidney Fs = Foot sinus A = Ampulla
  • 27. The transport of the blood to and from the organs occur through artery. They don’t have capillary veins and arterioles.
  • 28. Reproductive System Has separated sexes (dioecious). The differences between the sexes are not visible at shell colour or pattern.
  • 29. Male reproductive system  The male reproductive tract consist of the testis and vas deferens, the seminal vesicle and the prostate gland, the penial sac, the penis and the penis sheath.  During mating activities, this penial papilla bends towards the sperm groove in which the sperm is conducted.  The sperm is conveyed in the penial duct at the base of the penis.  On erection the penis comes out of the penial sac and is grasped around on the lower part by a muscular penial sheath from the mantle.
  • 30.
  • 31. Female Reproductive System The egg cells are stored in the ovaries. To produce an egg, the egg cell is brought to the receptaculum seminis. In this place, the fertilization of the egg cells takes place. The spermatozoa (sperm cells) of the male can survive for more than a month in the female, and several egg clutches can be fertilized with a single copulation. The eggs were deposited above the water line.