Pests of castor_Binomics_Identification_Dr.UPR.pdf
Algae Cell Morphology and Classification
1.
2. Algae are photosynthetic eukaryotic organisms.
They are commonly found in aquatic environments including
freshwater, marine and brackish water.
They are either motile or non-motile.
Some of the motile and non-motile algae may form a colony known as
Coenonbium.
Algae possess the usual eukaryotic structures - Golgi apparatus,
mitochondria, Endoplasmic reticulum and a nucleus.
The algae includes both the microscopic unicellular to macroscopic
multicellular organisms.
Most of the algae are autotrophic characterized by their ability to use
the carbon-di-oxide as a carbon source and light as an energy source.
They reproduce both sexually and asexually.
4. Algae are generally classified on the basis of the following
characteristics:
Chemical and physical feature of the cell walls
Nature and properties of pigments that contribute to
photosynthesis
Morphological charactersitics of cells and thalli.
Habitat
Flagella number and the location of their insertion in motile
cells.
Reproductive behaviour
5. One of the best known algal classification was proposed by Fritsch
who divided them into 11 classes(1945).
Classification is based on pigments, flagella and reserve food material.
Eleven classes proposed by Fritsch are as follows:
1. Chlorophyceae
2. Xanthophyceae
3. Chrysophyceae
4. Bacillariophyceae
5. Cryptophyceae
6. Dinophyceae
7. Chloromonadineae
8. Euglenineae
9. Phaeophyceae
10. Rhodophyceae
11. Myxophyceae.
6. 1980 - Lee used fine structural aspects of organelles such as
chloroplast, ER, flagellum, eyespot and nucleus to classify the algae
into 6 divisions and 15 classes.
The 6 divisions are : Cyanophyta, Glaucophyta, Chromophyta,
Rhododphyta, Chlorophyta and Charophyta.
Rosowski and Parker (1982) classified algae into 15 classes.
Van den Hoek (1995) classified algae into 11 divisions:
Cyanophyta
Prochlorophyta
Glaucophyta
Rhodophyta
Heterokontophyta
Haptophyta
Cryptophyta
Dinophyta
Euglenophyta
Chloroarachniophyta
Cholorphyta
7. Chlorophyta is the division of the green algae.
They are principally freshwater species.
They are also found in sea water, and many other terrestial
ecosystems.
This division contain around 500 genera and around 8000
species.
There are many single celled forms and many colonial types of
green algae.
Many unicellular green algae are motile by flagella action.
Colonial types occur as spheres, filaments or plates.
8. Some species have special structures called
holdfasts, which anchor them to submerged
objects or aquatic plants.
The cells of the chlorophycophyta have a well-
defined nucleus and usually a cell wall and the
chlorophyll and other pigments are in
chloroplasts as in higher plants.
Majority of the green algae contain one
chloroplasts per cell.
The cholorplasts contains chlorophyll a and b.
Green algae reproduce both sexually and
asexually.
The cholorplasts are green because they are not masked by any
accessory pigments. Because of this character, the members of
chlorophyta are called as green algae.
The chlorophyta has a characteristic set of acessory pigments such as
β-carotene and Xanthophylls.
9. Chlamydomonas:
It is a typical unicellular motile green algae widely distributed in
stagnant water, damp soils and even in snow as “Snow algae”.
(Chlamydomonas nivalis - found in snow).
They reproduce both sexually and asexually.
Model organism
10. Volvox:
Volvox is a colonial green algae.
Found in freshwater habitats.
It forms spherical colonies of up to 50,000 cells.
The individual cells are biflagellate and are morphologically similar to
that of Chlamydomonas.
11. Chlorella:
It is a green algae which is mainly found in freshwater habitats such as
ponds, ditches and lakes.
It is globular in shape.
It is non-motile in nature.
Generally used in supplemental food supply - SCP.
12. Ulothrix:
It is a filamentous green algae found in freshwater and marine water.
They are found in flowing streams.
They are attached to the twigs or stones by holdfasts at the bases of
the filaments.
It reproduces by means of the flagellated zoospores. Each Zoospore
may become attached to a solid object and develop in to a filament.
13. Spirogyra:
Also known as pond silk, water silk or blanket weed.
It is a filamentous green algae.
Named for the helical or spiral arrangement of the chloroplasts.
Spirogyra is very common in relatively clean eutrophic water,
developing slimy filamentous green masses.
14. Acetabularia:
The name, Acetabularia, derives from the Latin word acetabulum, a
broad, shallow cup used for dipping bread; the upturned cap of
Acetabularia resembles such a cup. For this reason, it is also
sometimes called mermaid's wineglass.
Each Acetabularia cell is composed of three segments: the "foot" or
base which contains the nucleus, the "stalk," and the "cap."
15. The members of this division
are red in colour. Hence they
are known as “Red Algae”.
Red algae are red because of
the presence of the pigment
phycoerythrin.
They are generally found in
frehwater or marine water.
The members of this division
are generally multicellular
though unicellualr forms are
known.
16. Some rhodophytes are also important in the formation of tropical reefs.
These reef-building rhodophytes are called coralline algae, because
they secrete a hard shell of carbonate around themselves, in much the
same way that corals do.
In many red algae, asexual reproduction takes place by non motile
spores called monospores.
Sexual reproduction is oogamous type involving specialised female
cells called Carpogonia and specialised male cells called spermatia.
Several red algae is of economic importance:
Carrageenan : It is extracted from the walls of several red algae. It is used
as a stabilizer or emulsifier in foods such as ice cream, custards and
evaporated milk.
Agar, a well-known solidifying agent used in the preparation of
microbiological media is extracted from Gelidium and Gracilaria.
17. Gelidium:
Gelidium is a genus of
thalloid red algae comprising
124 species.
G.amansii - Used in making agar.
Gracilaria:
Gracilaria is a genus of red
algae notable for its economic
importance in producing agar.
It is generally used as a food
source for humans and certain
fishes such as tangs.
18. The cryptomonads are a small
group of biflagellate organisms.
They have two unequal flagella,
which arise from the base of
the groove.
The cells are slipper shaped
and occur singly.
Some forms have a cellulose
wall while others are naked,
being surrounded by
plasmalemma with a thin
granular material on the
outside.
19. There are one or two plastids, with or without pyrenoids
per cell.
Food reserve is stored as a true starch as well as oil.
The longer flagellum bears two rows of mastigonemes,
while the shorter flagellum bears a single row of
mastigonemes.
The choloroplasts contain chlorophylls a and c, but their
colour is masked by a variety of accessory pigments
which occur in different proportions.
The pigments include phycocyanin, phycoerythrin, α-
carotene and the Xanthophylls.
Reproduction is either by means of longitudinal cell
division or the formation of zoospores or cysts.
20.
21. Cryptomonas:
It is a very important member of the
cryptophyta division.
They are widely distributed in freshwater
habitats.
Cryptomonas is a dimorphic genus,
meaning it could be either protozoan
(Cryptomondida) or alga
(Cryptophyceae).
The cells are usually brownish in color, and have a slit-like furrow
at the anterior.
They are not known to produce any toxins and are used to feed
small zooplankton, which is the food source for small fish in fish
farming.
Currently there are 26 species of Cryptomonas.
22. The members of this algal division are commonly called as
euglenoids, because the best known genus of the group is Euglena.
Most of the euglenophyta are freshwater organisms, but some are
marine.
There are around 800 species of euglenophytes.
They are unicellular flagellates.
There are almost always two flagella, but one of these is often very
short.
The members lack a cell wall but normally are surrounded by an
outer layer of pellicle composed of proteins and lipids.
The euglenoid algae contain chlorophylls a and b like the green
algae and typically appear green because the chlorophylls are not
masked by any accessory pigments.
23. Of the accessory pigments that are present, the most important are
β-carotene, neoxanthin and diadinoxanthin, although echinenone,
diatoxanthin and zeaxanthin are also found.
This group is known to contain carbohydrate paramylon.
Nutrition:
A) Photosynthetic:
They can supplement photosynthesis by taking up the organic
compounds. Eg: Euglena.
B) Heterotrophic:
There are also many colourless euglenophytes which are completely
dependent on heterotrophic nutrition. Eg: Astasia.
C) Phagotrophic:
Few euglenophytes are phagotrophic and possess a special apparatus
for capturing the prey and a cytostome for ingesting them. Eg:
Peranema.
24.
25. Euglena:
They are both plant-like and animal-like.
They are unicellular.
They are motile by means of flagellum.
They commonly occur in freshwater and salt water.
There is no cell wall containing cellulose.
The outer membrane is an organised periplast.
Certain species develop a prominent stigma or eye spot.
Contractile vacuoles are also present.
Most species of Euglena have photosynthesizing chloroplasts
within the body of the cell, which enable them to feed by
autotrophy, like plants.
26. In low moisture conditions, or when food is scarce, Euglena forms
a protective wall around itself and lies dormant as a resting cyst
until environmental conditions improve.
All Euglenoids have two flagella rooted in basal bodies located in a
small reservoir at the front of the cell.
In Euglena, one flagellum is very short, and does not protrude from
the cell, while the other is relatively long, and often easily visible
with light microscopy.
27. It is a large group of algae
commonly found in the
freshwater.
Species of chrysophycophyta
are predominantly flagellates.
Some are amoeboid with the
psudopodial extensions of the
protoplasm.
Most forms are unicellular, but
some form colonies.
They differ from the green algae
in the nature of their pigments,
in storing reserve food as oil or
chrysolaminarin rather than
starch.
28. Their characteristic colour is due to the masking of their
chlorophyll by brown pigments.
Reproduction is commonly by binnary fission but occasionally
isogamous.
Ochromonas:
It is an unicellular genus with
unequal flagellation.
They are either spherical or
cylindrical in shape.
They commonly appear in
freshwater, brackish or marine
waters.
Some species (especially
Ochromonas danica) are important
in physiological and molecular
studies.
29. Chrysamoeba:
They are unicellular flagellates.
The flagella are unequal in length.
There are two distinctive plastids in the photosynthetic forms.
It is an amoeboid cell with a number of long rhizopodia radiating
from the central mass of protoplasm.