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Detailed study of pila

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Detailed study of pila

  1. 1. GUIDED BY – Dr. Poonam Sharma Made By – Kuldeep Gauliya
  4. 4. SENSE ORGANS OF Pila • Pila have special organs of sense. • It consists of these organs :- (A) A single osphradium (B) Paired eyes (C) Statocyst (D) Tentacles
  5. 5. A. OSPHRADIUM • Osphradium hangs from the mentle near the pseudodipodium. • It is oval with 22 to 28 fleshy leaflets arranged on the sides of central axis.
  6. 6. B. STATOCYST • Located in the foot near each pedal ganglion like a statocyst in depression. • It is a round capsule lined with epithilial cells and surrounded by connective tissue. • The statocyst receive the nerves from pedal & cerebral ganglia, they are the organs of equilibirium and regulate the position of the snail.
  7. 7. C. EYES • An eyes is an oval capsule, its wall is the ratina made of pigmented sensory cells, it is continued in front as a thin, non pigmented, transparent cornea. • An opotic nerve innervates the ratinal cells. • Eyes are sensory to light.
  8. 8. D. TENTACLES • The tentacles and foot are liberally supplied with nerves, they are sensory to contacts, tentacles contains both tactile and chemoreceptor cells and probably gustatory also. • The first pair of tentacles are olfactory.
  9. 9. REPRODUCTIVE SYSTEM • In Pila sexes are separate, i.e. dioecious and there is a definate sexual dimorphism. • The shell of the male is smaller in size and less swollen then the female. • There is a well develop copulatory organ in the male but it is quit redimentary in the female.
  10. 10. MALE REPRODUCTIVE SYSTEM• The male reproductive system consist of 1. Testis with fine vasa efferentia, 2. Vas deferens with the vesicula seminalis and the terminal glandular part of the vas deferens, 3. panis with its sheath and, 4. Hypobranchial glands.
  11. 11. TESTIS • It is the flate plate like whitish structure,more or less triangular in outline situated in the upper part of the first whorls of the shell. • The cream colored testis is easily distinguished from the from the digestive gland which is brownish or dirty green. • Minute ducts the vasa efferentia lead downwards from the different
  12. 12. parts of the testis and may unite with one another before opening into vas deferens.
  13. 13. VAS DEFERENS • From the posterior end of testis arises a thin vas deferens. • It consist of three distincts parts:- 1. Proximal thin tubular portion leading from the testis. 2. Vesicula seminalis, 3. Thick glandular portion which opens into the mantal cavity near the anul opening.
  14. 14. # The vesicula seminalis open on the left side into the terminal glandular part of the vas deferens. # In the mantal- cavity, the vas deferens lies closely attached to the left side of the rectum and the ends in a prominent claw – shaped structure the genital papilla having the male genital aperture a littile behind the anus.
  15. 15. PENIS SHEATH AND PENIS • The edge of the mantal bears on its inner surface a thick glandular flap of a yellowish color. • The flap is attached on its right-side but is free on its left ; edge slightly rolled in to from s spout – like sheath. • The penis is capable of extension.
  16. 16. HYPOBRANCHIAL GLAND • At the base of the penis sheath is an oval hypobranchial gland. • It consist of of tall cells containing small nuclie.
  17. 17. SPERMATOGENESIS # The spermatogenesis of Pila globosa has been worked out by Sharma, G.P, Gupta, BrijLal, and Mital,O.P. (1959) and they have reported that the spermatozoa of Pila are of two kinds :- (1) Eupyrene sperm (2) Oligopyrene sperm
  18. 18. 1. EUPYRENE SPERMS • The eupyrene sperms are hair like having and elpongated spiraly twisted nucleus with a small conical acrosome in front, and a mitochondrialmiddle piece behind, followed by the end piece in the form of a long vibrating tail. • These sperms moove actively forward in the spiral course, measure about 25 µ in length and 1.2 µ in breadth and they can only fertilise the eggs.
  19. 19. OLIGOPYRENE SPERM • The oligopyrene sperm on the other hand, have a very sluggish and serpent – like movement. • Ther acrosome is poorly developed, the nucleus is elongated, broad curved but but spirally coiled ; the middle piece is short and the no. of axial filaments varies from 4 – 8. • These are probably having some secondary function.
  20. 20. FEMALE REPRODUCTIVE SYSTEM The female reproductive system is consists of 1. Ovary with numerous minute ducts, 2. main oviduct, 3. receptaculum seminis, 4. uterus, 5.Vagina and, 6. Hypobranchial gland.
  21. 21. 1. OVARY • The ovary in the female lies in the same position as the testis in the male but it is not so extensive. • Ovary is much branched structure of a light orange color which become Darker in fully matured individuals. • The branches of ovary consist of single layered acini which are more or less flask – shaped.222
  22. 22. 2. OVIDUCT • The narrow and transport oviduct originates from about the middle of the ovary. • It runs anteriorly just below the skin along the inner margin of the digestive gland. • Near the renal organ it turns downward and then upwards to enter the receptaculum seminis.
  23. 23. RECEPTACULUM SEMINIS • It is a bean shaped structure. • A thin walled pouch arises directly from the wall of the uterus and is called the pouch of the receptaculum.
  24. 24. 4. UTERUS • It is a large pear shaped structure, deep yellow in color. • It lies inside the inside the body whorl below the intestine and the right of the renal chambers. • The apex of the uterus points forwards and is continued as the vagina, while its basal portion is broad and rounded and is connected on its outer side with the receptaculum seminis
  25. 25. 4. VAGINA • The vagina is the white or cream colored, band like structure laying immediately beneath the skin. • It extends from the uterus to the upper end of columellar muscle.The vagina enters the mantle cavity at its right posterior corner and continues farwads to the female genital apereture situated on the smaal papilla, a little behind the anus.
  26. 26. 6. HYPOBRANCHIAL GLAND • The hypobranchial cavity of the female is poorly developed. • There is a rudimentary glandular thickning in the area of the hypobranchial gland.
  27. 27. 7. COPULATORY APPARATUS • The female is the rudimentary penis laying beneath the glandular fold at the edge of the mantal. • There is no trace of the foldings of the penis sheath.
  28. 28. FERTILESATION • Eggs are fertilised in the uterus and oviposition starts a day or two later. • The fertilised eggs are laid in masses of 200 to 800 in moist earth near pond and lakes.
  29. 29. DEVELOPMENT • In development mollusca passes through two larval stages, there is a trochosphere larva which soon grows into veliger larva. • A free swimming trochosphere is found only in some ptimitive gastropodes, such as Diotocardia, but in all others the trochosphere stage is reduced and passed within the egg.