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N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and
                  Applications (IJERA) ISSN: 2248-9622 www.ijera.com
                   Vol. 2, Issue 5, September- October 2012, pp.521-523
     Diversity and abundance of Dragonflies and Damselflies Of
     Chatri Lake Region, in Pohara – Malkhed Reserve Forest,
                   Amravati, Maharashtra (India)
                     N.A. Manwar*1, P.P. Rathod2 and I. Ahmad Raja1
                       1
                       Shri Shivaji College of Arts, Commerce and Science, Akola - 444001
                2
                    Government Vidarbha Institute of Science and Humanities, Amravati - 444604

Abstract
        The objectives of the present study were          and Damselflies of Chatri Lake. The present work
to explore the Diversity and abundance of                 was, therefore, undertaken to make the study on
Dragonflies and Damselflies (Class Insecta,               Diversity and abundance of Dragonflies and
phylum Arthopoda) of Chatri Lake region.                  Damselflies of Chatri Lake region.
Diversity and abundance of the Chatri Lake
region was investigated. Total 22 species of
dragonflies and damselflies belonging to four             2. Study area:
families were recorded. In our study the most                       Chatri Lake is situated in Pohara – Malkhed
abundant family was Lebellulidae followed by              Reserve Forest, Amravati and it is also nearer to
coenogrionidae, while family gomphidae and                Amravati City. The Chatri Lake lies between
Platycenemididae      was      least    abundant.         (20053’42.6”N & 77046’66.2”E, 372m), covers an
Lebellulidae family represents 11 species;                area of 111.231934 m2. Chatri Lake is an artificial
coenogrionidae represents 8 species while                 lake and was built in 1888. It is situated on the
gomphidae and Platycenemididae represents two             Malkhed Railway Road, 1 km from Dasturnagar
and one species respectively. We observed that the        Square; the reservoir has its base built on a small
station 1 is most diverse in Dragonfly and                spring named Kali Nadi. It is a small reservoir built
Damselfly fauna.                                          with the intention to supply drinking water to
                                                          Amravati City, but now Amravati City gets water
Key words: Dragonfly and damselflies, Chatri              from Upper Wardha Dam. A small garden and
Lake, Pohara – Malkhed Reserve Forest, Amravati.          boating facility is also available, because of beautiful
                                                          garden and boating facility many tourist visits this
1. Introduction:                                          place. Human activities, fishing and cattle grazing is
          The order Odonata is one of the most            also seen on this Lake.
popular insect groups. Dragonflies and damselflies        For better study we have divided this area into three
are amongst the most attractive of creatures on earth.    different stations. These stations are
According to Silsby, (2001), eight super families, 29     1)        STATION 1 – Garden site
families and some 58 sub-families of dragonflies for      2)        STATION 2 – Most contaminated cattle
approximately 600 genera and 6000 named species                     grazing site
have so far been described all over the world. India is   3)        STATION 3 – Fish collection site
also highly diverse with more than 500 known
species of Odonata (Subramanian, 2005). They are
denizens of many aquatic ecosystems and their
distribution covers a great deal of continuum from
temporary to permanent water bodies (Corbet 1999;
Johansson & Suhling 2004). In the temperate regions
of the world, dragonflies are frequently used as
indicators of environmental health. Their aquatic
larvae constitute a natural biological control over
mosquito larvae and thus help to control several
epidemic diseases like malaria, dengue, filaria etc.
(Mitra, 2002). In India Odonata status and give us
valuable insight about ecosystem health, especially of
wetland. They are among the dominant invertebrates
predators in any ecosystem. Being predators both at
larval and adult stages, they play significant role in
the food chain of the forest ecosystem (Vashishth
et.al, 2002). They tend to reside in flowing as well as
in standing waters (Corbet, 1962). There is, however
no report on Diversity and abundance of Dragonflies


                                                                                                  521 | P a g e
N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and
                  Applications (IJERA) ISSN: 2248-9622 www.ijera.com
                   Vol. 2, Issue 5, September- October 2012, pp.521-523
                                                          during morning and evening times in all stations.
                                                          Collection of specimens was done with the help of
                                                          specially design insect net. Species were
                                                          photographed and identified in their natural habitats,
                                                          but in few cases when it is difficult to assess, then it
                                                          is collected for further identification. Standard
                                                          methods were used for collection and observations of
                                                          species. Species were identified with the help of
                                                          Fauna of British India of Fraser and field guide of
                                                          Subramanian.

                                                          4. Results:
                                                          In our study 22 species of dragonflies and damselflies
                                                          to four families and 17 genera were recorded. We
                                                          observed that out of total species recorded 50% are of
                                                          Lebellulidae family followed by coenogrionidae
                                                          family with 36% species. Gomphidae and
                                                          Platycenemididae family showed less species
                                                          diversity and represented by only two and one
                                                          species respectively (fig. 2). Station wise status of
                                                          each species was investigated. Station 1 was very
                                                          much diverse and almost all species except few are
                                                          reported in this area. Station 2 was very much
                                                          contaminated and has very few species at that site,
                                                          but Brachythemis contaminata was seen most
                                                          abundant. Station 3 was also contaminated but it is
                                                          less contaminated as compared to station 2 and has
Fig. 1 Study area (Chatri Lake, Amravati)
                                                          some more species diversity than station 2.
3.      Materials and methods:
The present study was carried out monthly during the
year June-Nov 2011. Observations were carried out

Table 1 – Name of species and their status in different station (+ indicate presence while – indicate absence)
SR.N     NAME OF SPECIES                                              STATION 1 STATION 2              STATION 3
O.
1        Orthetrum chrysis (Selys, 1891)                              -              -                 +
2        Orthetrum sabina (Drury, 1770)                               +              -                 +
3        Orthetrum glaucaum (Brauer, 1865)                            +              -                 -
4        Orthetrum prinosum (Rambur, 1842)                            +              -                 -
5        Rhyothemis variegata (Linnaeus, 1763)                        +              -                 -
6        Brachythemis contaminata (Fabricius, 1793)                   +              +                 +
7        Anthriamanta brevipennis (Rambur, 1842)                      +              -                 -
8        Trithemis festiva (Rambur, 1842)                             -              +                 -
9        Trithemis pallidinervis (Kirby , 1889)                       +              +                 -
10       Diplocodes trivialis (Rambur, 1842)                          +              -                 +
11       Crocothemis servilia (Drury, 1770)                           -              -                 +
12       Ictinogomphus rapax (Rambur, 1842)                           +              +                 +
13       Paragomphus lineatus (Selys, 1850)                           +              -                 +
14       Acisoma panorpoides (Rambur, 1842)                           +              -                 -
15       Pseudagrion rubriceps (Selys, 1876)                          +              -                 -
16       Pseudagrion microcephalum (Rambur, 1842)                     +              +                 -
17       Aciagrion pallidum (Selys, 1891)                             +              -                 -
18       Ischnura arora (Brauer, 1865)                                +              -                 +
19       Rhodischnura nursei (Morton, 1907)                           +              +                 -
20       Ceriagrion coromandelianum (Fabricius, 1798)                 +              -                 -
21       Mortonagrion varralli (Fraser, 1920)                         -              +                 -
22       Copera marginipes (Rambur, 1842)                             +              -                 -




                                                                                                  522 | P a g e
N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and
                    Applications (IJERA) ISSN: 2248-9622 www.ijera.com
                     Vol. 2, Issue 5, September- October 2012, pp.521-523
Family: Libellulidae                              7) Mortonagrion varralli (Fraser, 1920) – Found
1) Orthetrum chrysis (Selys, 1891) – Found                  among bushes close to marshes.
    commonly perched arraund marshes, ponds.             8) Rhodischnura nursei (Morton, 1907) – In
2) Orthetrum sabina (Drury, 1770) – A common                marshes and ponds.
    dragonfly of garden and fields.
3) Orthetrum glaucaum (Brauer, 1865) –                   Family: Platycenemididae
    Commonly found in canals.                            1) Copera marginipes (Rambur, 1842) – Found
4) Orthetrum prinosum (Rambur, 1842) – A                    along ponds, canals and streams. Fly along the
    common dragonfly of ponds, ditches, wells,              ground level.
    tanks, rivers.
5) Rhyothemis variegata (Linnaeus, 1763) –
    Predominant dragonfly of pond, marshes.
6) Anthriamanta brevipennis (Rambur, 1842) –
    They have adapted to urban environment and                                             Libellulidae
    could be seen in garden ponds too.                                                     (50%)
7) Trithemis festiva (Rambur, 1842) – Very                                                 Gomphidae
    common in canals, usually perches on boulders                                          (9%)
    and aquatic plants.
8) Trithemis pallidinervis (Kirby , 1889) – A                                              Coenagrionid
    dragonfly partial to the marshes and weedy                                             ae (36%)
    ponds. Usually perch on bare tips of shrubs. The
    long legs are very noticeable at this time.                                            Platycenemi
9) Diplocodes trivialis (Rambur, 1842) – common                                            didae (5%)
    dragonflies in gardens, fields etc. This dragonfly
    usually perches on ground and rarely flies above
    1m.
10) Crocothemis servilia (Drury, 1770) – One of the
                                                         Fig.1 - Distribution of Odonata families from Chatri
    commonest red dragonflies. This dragonfly
                                                         Lake region.
    perches on aquatic weeds and chases any passing
    by dragonflies.
11) Brachythemis contaminata (Fabricius, 1793) – A
                                                         5. Discussion:
                                                                   So far, the occurrence of 22 species of
    dragonfly of polluted waters.
                                                         dragonflies and damselflies was reported under four
Family: Gomphidae                                        families from Amravati. Among these Brachythemis
1) Ictinogomphus rapax (Rambur, 1842) –
                                                         contaminata was abundant and hence, indicate that
    Common dragonfly usually perches on a bare
                                                         this Lake may be contaminated. Station 1 is most and
    twig facing the water
                                                         Station 2 is list diverse in dragonfly and damselfly
body.
                                                         diversity. From this study we conclude that our study
2) Paragomphus lineatus (Selys, 1850) –
                                                         area is very much diverse in dragonfly and damselfly
    Commonly found near streams, rivers, ponds and
                                                         fauna.
    lakes.
                                                         Acknowledgement:
Family: Coenagrionidae                                            Authors are thankful to The Principal of
1) Ischnura arora (Brauer, 1865) – Among marshes
                                                         college Shree Shivaji College of Arts, Commerce and
   on the banks of ponds, canals and rivers.
                                                         Science, Akola, for their permission and guidance to
2) Pseudagrion rubriceps (Selys, 1876) – Seen in
                                                         such study.
   small groups of 3-4 individuals.
3) Aciagrion pallidum (Selys, 1891) - Common in
   marshes and ponds, flying or perched about 1-2
                                                         References:
   feet above ground.                                           P. S. Corbet, (1962) Biology of dragonflies.
4) Ceriagrion coromandelianum (Fabricius, 1798)                  Witherby, Londan, 247 pp.
   – One of the commonest damselfly of central                  P. S. Corbet, (1999) Dragonflies: Behaviour
   India, found along banks of pond, garden tanks,               and Ecology of Odonata.829pp
   rivers and canals. Also found away from the                  F. C. Fraser, The fauna of British India
   water bodies.                                                 including Ceylon and Burma, Odonata, vol.I
5) Acisoma panorpoides (Rambur, 1842) – A                        Taylor     & Francis Ltd.,London.423pp
   species closely associated with water.                        .(1933) .
6) Pseudagrion microcephalum (Rambur, 1842) –                   F. C. Fraser, The fauna of British India
   Found commonly among vegetation covered                       including Ceylon and Burma, Odonata, vol.I
   banks of ponds, canals and rivers.                            Taylor     & Francis Ltd.,London.398pp
                                                                 .(1934) .



                                                                                               523 | P a g e
N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and
                  Applications (IJERA) ISSN: 2248-9622 www.ijera.com
                   Vol. 2, Issue 5, September- October 2012, pp.521-523
      F. C. Fraser, The fauna of British India
       including Ceylon and Burma, Odonata, vol.I
       Taylor      & Francis Ltd.,London.461pp
       .(1936) .
      F. Johansson, & F. Suhling, Behaviour and
       growth of dragonfly larvae along a
       permanent to temporary water habitat
       gradient. Ecological Entomology 29: 196-
       202 (2004).
      A. Mitra, Dragonfly (Odonata: Insecta)
       Fauna of Trashigang Dzongkhag, Eastern
       Bhutan. In: Gyeltshen, T. & Sadruddin (ed.)
       “Environment and Life Support Systems of
       the Bhutan Himalaya, Vol. I”, Sherubtse
       College, Kanglung, Bhutan: 40-70 (2002).
      J. Silsby, Dragonflies of the World. Natural
       History Museum in association with CSIRO
       Publishing,UK and Europe (2001).
      K. A. Subramanian, Dragonflies and
       damselflies of Penisular India - A field
       guide. Project Lifescape. Indian Academy of
       Sciences, Banglore, India. Pp.118 (2005).
      N. Vashishth, P.C. Joshi & A. Singh,
       Odonata community dynamics in Rajaji
       National Park, India Fraseria 7: 21-25
       (2002)




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  • 1. N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.521-523 Diversity and abundance of Dragonflies and Damselflies Of Chatri Lake Region, in Pohara – Malkhed Reserve Forest, Amravati, Maharashtra (India) N.A. Manwar*1, P.P. Rathod2 and I. Ahmad Raja1 1 Shri Shivaji College of Arts, Commerce and Science, Akola - 444001 2 Government Vidarbha Institute of Science and Humanities, Amravati - 444604 Abstract The objectives of the present study were and Damselflies of Chatri Lake. The present work to explore the Diversity and abundance of was, therefore, undertaken to make the study on Dragonflies and Damselflies (Class Insecta, Diversity and abundance of Dragonflies and phylum Arthopoda) of Chatri Lake region. Damselflies of Chatri Lake region. Diversity and abundance of the Chatri Lake region was investigated. Total 22 species of dragonflies and damselflies belonging to four 2. Study area: families were recorded. In our study the most Chatri Lake is situated in Pohara – Malkhed abundant family was Lebellulidae followed by Reserve Forest, Amravati and it is also nearer to coenogrionidae, while family gomphidae and Amravati City. The Chatri Lake lies between Platycenemididae was least abundant. (20053’42.6”N & 77046’66.2”E, 372m), covers an Lebellulidae family represents 11 species; area of 111.231934 m2. Chatri Lake is an artificial coenogrionidae represents 8 species while lake and was built in 1888. It is situated on the gomphidae and Platycenemididae represents two Malkhed Railway Road, 1 km from Dasturnagar and one species respectively. We observed that the Square; the reservoir has its base built on a small station 1 is most diverse in Dragonfly and spring named Kali Nadi. It is a small reservoir built Damselfly fauna. with the intention to supply drinking water to Amravati City, but now Amravati City gets water Key words: Dragonfly and damselflies, Chatri from Upper Wardha Dam. A small garden and Lake, Pohara – Malkhed Reserve Forest, Amravati. boating facility is also available, because of beautiful garden and boating facility many tourist visits this 1. Introduction: place. Human activities, fishing and cattle grazing is The order Odonata is one of the most also seen on this Lake. popular insect groups. Dragonflies and damselflies For better study we have divided this area into three are amongst the most attractive of creatures on earth. different stations. These stations are According to Silsby, (2001), eight super families, 29 1) STATION 1 – Garden site families and some 58 sub-families of dragonflies for 2) STATION 2 – Most contaminated cattle approximately 600 genera and 6000 named species grazing site have so far been described all over the world. India is 3) STATION 3 – Fish collection site also highly diverse with more than 500 known species of Odonata (Subramanian, 2005). They are denizens of many aquatic ecosystems and their distribution covers a great deal of continuum from temporary to permanent water bodies (Corbet 1999; Johansson & Suhling 2004). In the temperate regions of the world, dragonflies are frequently used as indicators of environmental health. Their aquatic larvae constitute a natural biological control over mosquito larvae and thus help to control several epidemic diseases like malaria, dengue, filaria etc. (Mitra, 2002). In India Odonata status and give us valuable insight about ecosystem health, especially of wetland. They are among the dominant invertebrates predators in any ecosystem. Being predators both at larval and adult stages, they play significant role in the food chain of the forest ecosystem (Vashishth et.al, 2002). They tend to reside in flowing as well as in standing waters (Corbet, 1962). There is, however no report on Diversity and abundance of Dragonflies 521 | P a g e
  • 2. N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.521-523 during morning and evening times in all stations. Collection of specimens was done with the help of specially design insect net. Species were photographed and identified in their natural habitats, but in few cases when it is difficult to assess, then it is collected for further identification. Standard methods were used for collection and observations of species. Species were identified with the help of Fauna of British India of Fraser and field guide of Subramanian. 4. Results: In our study 22 species of dragonflies and damselflies to four families and 17 genera were recorded. We observed that out of total species recorded 50% are of Lebellulidae family followed by coenogrionidae family with 36% species. Gomphidae and Platycenemididae family showed less species diversity and represented by only two and one species respectively (fig. 2). Station wise status of each species was investigated. Station 1 was very much diverse and almost all species except few are reported in this area. Station 2 was very much contaminated and has very few species at that site, but Brachythemis contaminata was seen most abundant. Station 3 was also contaminated but it is less contaminated as compared to station 2 and has Fig. 1 Study area (Chatri Lake, Amravati) some more species diversity than station 2. 3. Materials and methods: The present study was carried out monthly during the year June-Nov 2011. Observations were carried out Table 1 – Name of species and their status in different station (+ indicate presence while – indicate absence) SR.N NAME OF SPECIES STATION 1 STATION 2 STATION 3 O. 1 Orthetrum chrysis (Selys, 1891) - - + 2 Orthetrum sabina (Drury, 1770) + - + 3 Orthetrum glaucaum (Brauer, 1865) + - - 4 Orthetrum prinosum (Rambur, 1842) + - - 5 Rhyothemis variegata (Linnaeus, 1763) + - - 6 Brachythemis contaminata (Fabricius, 1793) + + + 7 Anthriamanta brevipennis (Rambur, 1842) + - - 8 Trithemis festiva (Rambur, 1842) - + - 9 Trithemis pallidinervis (Kirby , 1889) + + - 10 Diplocodes trivialis (Rambur, 1842) + - + 11 Crocothemis servilia (Drury, 1770) - - + 12 Ictinogomphus rapax (Rambur, 1842) + + + 13 Paragomphus lineatus (Selys, 1850) + - + 14 Acisoma panorpoides (Rambur, 1842) + - - 15 Pseudagrion rubriceps (Selys, 1876) + - - 16 Pseudagrion microcephalum (Rambur, 1842) + + - 17 Aciagrion pallidum (Selys, 1891) + - - 18 Ischnura arora (Brauer, 1865) + - + 19 Rhodischnura nursei (Morton, 1907) + + - 20 Ceriagrion coromandelianum (Fabricius, 1798) + - - 21 Mortonagrion varralli (Fraser, 1920) - + - 22 Copera marginipes (Rambur, 1842) + - - 522 | P a g e
  • 3. N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.521-523 Family: Libellulidae 7) Mortonagrion varralli (Fraser, 1920) – Found 1) Orthetrum chrysis (Selys, 1891) – Found among bushes close to marshes. commonly perched arraund marshes, ponds. 8) Rhodischnura nursei (Morton, 1907) – In 2) Orthetrum sabina (Drury, 1770) – A common marshes and ponds. dragonfly of garden and fields. 3) Orthetrum glaucaum (Brauer, 1865) – Family: Platycenemididae Commonly found in canals. 1) Copera marginipes (Rambur, 1842) – Found 4) Orthetrum prinosum (Rambur, 1842) – A along ponds, canals and streams. Fly along the common dragonfly of ponds, ditches, wells, ground level. tanks, rivers. 5) Rhyothemis variegata (Linnaeus, 1763) – Predominant dragonfly of pond, marshes. 6) Anthriamanta brevipennis (Rambur, 1842) – They have adapted to urban environment and Libellulidae could be seen in garden ponds too. (50%) 7) Trithemis festiva (Rambur, 1842) – Very Gomphidae common in canals, usually perches on boulders (9%) and aquatic plants. 8) Trithemis pallidinervis (Kirby , 1889) – A Coenagrionid dragonfly partial to the marshes and weedy ae (36%) ponds. Usually perch on bare tips of shrubs. The long legs are very noticeable at this time. Platycenemi 9) Diplocodes trivialis (Rambur, 1842) – common didae (5%) dragonflies in gardens, fields etc. This dragonfly usually perches on ground and rarely flies above 1m. 10) Crocothemis servilia (Drury, 1770) – One of the Fig.1 - Distribution of Odonata families from Chatri commonest red dragonflies. This dragonfly Lake region. perches on aquatic weeds and chases any passing by dragonflies. 11) Brachythemis contaminata (Fabricius, 1793) – A 5. Discussion: So far, the occurrence of 22 species of dragonfly of polluted waters. dragonflies and damselflies was reported under four Family: Gomphidae families from Amravati. Among these Brachythemis 1) Ictinogomphus rapax (Rambur, 1842) – contaminata was abundant and hence, indicate that Common dragonfly usually perches on a bare this Lake may be contaminated. Station 1 is most and twig facing the water Station 2 is list diverse in dragonfly and damselfly body. diversity. From this study we conclude that our study 2) Paragomphus lineatus (Selys, 1850) – area is very much diverse in dragonfly and damselfly Commonly found near streams, rivers, ponds and fauna. lakes. Acknowledgement: Family: Coenagrionidae Authors are thankful to The Principal of 1) Ischnura arora (Brauer, 1865) – Among marshes college Shree Shivaji College of Arts, Commerce and on the banks of ponds, canals and rivers. Science, Akola, for their permission and guidance to 2) Pseudagrion rubriceps (Selys, 1876) – Seen in such study. small groups of 3-4 individuals. 3) Aciagrion pallidum (Selys, 1891) - Common in marshes and ponds, flying or perched about 1-2 References: feet above ground.  P. S. Corbet, (1962) Biology of dragonflies. 4) Ceriagrion coromandelianum (Fabricius, 1798) Witherby, Londan, 247 pp. – One of the commonest damselfly of central  P. S. Corbet, (1999) Dragonflies: Behaviour India, found along banks of pond, garden tanks, and Ecology of Odonata.829pp rivers and canals. Also found away from the  F. C. Fraser, The fauna of British India water bodies. including Ceylon and Burma, Odonata, vol.I 5) Acisoma panorpoides (Rambur, 1842) – A Taylor & Francis Ltd.,London.423pp species closely associated with water. .(1933) . 6) Pseudagrion microcephalum (Rambur, 1842) –  F. C. Fraser, The fauna of British India Found commonly among vegetation covered including Ceylon and Burma, Odonata, vol.I banks of ponds, canals and rivers. Taylor & Francis Ltd.,London.398pp .(1934) . 523 | P a g e
  • 4. N.A. Manwar, P.P. Rathod, I. Ahmad Raja / International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.521-523  F. C. Fraser, The fauna of British India including Ceylon and Burma, Odonata, vol.I Taylor & Francis Ltd.,London.461pp .(1936) .  F. Johansson, & F. Suhling, Behaviour and growth of dragonfly larvae along a permanent to temporary water habitat gradient. Ecological Entomology 29: 196- 202 (2004).  A. Mitra, Dragonfly (Odonata: Insecta) Fauna of Trashigang Dzongkhag, Eastern Bhutan. In: Gyeltshen, T. & Sadruddin (ed.) “Environment and Life Support Systems of the Bhutan Himalaya, Vol. I”, Sherubtse College, Kanglung, Bhutan: 40-70 (2002).  J. Silsby, Dragonflies of the World. Natural History Museum in association with CSIRO Publishing,UK and Europe (2001).  K. A. Subramanian, Dragonflies and damselflies of Penisular India - A field guide. Project Lifescape. Indian Academy of Sciences, Banglore, India. Pp.118 (2005).  N. Vashishth, P.C. Joshi & A. Singh, Odonata community dynamics in Rajaji National Park, India Fraseria 7: 21-25 (2002) 524 | P a g e