1. A novel method for time-binning
rates of continuous character
evolution on a phylogeny
Graeme T. Lloyd and Matt Friedman

2. Coming soon…
Fossilworks Intensive Workshop in Analytical
Paleobiology
June-July 2014
University of Macquarie, Sydney
fossilworks.org

3. Acknowledgements
Chris
Organ
Graham
Slater

4. Phylogeny in macroevolution

5. Variable-rate models
Trait
Early Burst
Late Burst
Time

6. Rate
Continuous rates
Time

7. Rate
Continuous rates
Time

8. Continuous rates
Raia et al. 2013

9. Guts
Harmon et al. 2008

10. Method

11. Method

12. Method

13. Method

14. Method
S1
S2
1
S3

15. Data
Anderson et al. 2013

16. Data
Anderson et al. 2013

17. “Equal” time-scaling
Share time with preceding
(non-zero length) branch
Traditional approach first
τ8
τ8
τ7
τ7
τ6
τ6
τ5
τ5
root age
Ruta et al. 2006; Brusatte et al. 2008

18. Probabilistic time-scaling
τ6
τ5
τ4
τ3
τ2
τ1
τ0
Hedman 2010

19. Probabilistic time-scaling
τ6
τ5
τ4
τ3
τ2
τ1
τ0
Age (Ma)
τ0
Hedman 2010

20. “Equal” time-tree

21. Probabilistic time-tree

22. Rates
“Equal” timetree

23. Rates
“Equal”
timetree

24. Rates
“Equal” timetree
Probablistic timetree

25. “Equal” rates with error

26. Probabilistic rates with error

27. Model complexity
S1
S2

28. Change in mode
Ornstein-Uhlenbeck
Brownian
Slater 2013

29. Conclusions

30. Conclusions

31. Conclusions

32. Conclusions