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New concepts in maintenance of plant breeding promises and prospects
NEW CONCEPTS IN
MAINTENANCE OF
PLANT BREEDING
PROMISES AND
PROSPECTS
SUBMITTED TO
Dr. M.REDDI SEKHAR
PRESENTED BY
ZUBY GOHAR ANSARI
TAM/14/26
 NEW CONCEPTS IN WHOLE PLANT GENETICS
 In the excitement surrounding accomplishments in
the manipulation of tissues, cells, and genes over
the past ten years, one might get the impression
that plant breeding research at the whole plant and
population levels has been fixed away and
forgotten.
 On the contrary, several exciting developments in
breeding technology have surfaced during this
same period.
 One premise upon which the need for cellular and
molecular approaches has been based is that plant
breeders have exhausted existing pools of genetic
variability, and progress has consequently
diminished.
 There is little evidence to substantiate the existence
of such limitations for most crops. Moreover, it is
premature to conclude that sexually accessible
variability has been depleted when the existing
collections are rarely, if ever, comprehensive.
 New variability, for example, in soybean and rice
introductions (among others) from the People’s
Republic of China will more likely sustain
performance increases in the foreseeable future
than will new cellular/molecular approaches.
 The idea of expanding the pool of accessible genes
by overcoming natural isolating mechanisms
between species remains a possibility; hybrids
between species can be synthesized and
chromosomally altered to produce dramatic
economic gains, as with strawberries; it is now
possible to sexually engineer wheat lines containing
desired genes from related species simply by
adding or substituting whole chromosomes.
 In other research, crosses between the cultivated
tomato and a salt-tolerant wild species have
produced breeding lines with some salt tolerance.
 Preliminary observations suggest that cold
tolerance is another possibility.
 To date, 14 genes for disease resistance have been
transferred from wild species into cultivated tomato
varieties.
 Cell culture of interspecific hybrids has been
successfully applied to facilitate the flow of genes
among species related to tomato, barley, and
cabbage.
 Altogether, exploitation of genes from wild
germplasm sources has barely begun.
New concepts in maintenance of plant breeding promises and prospects
 A major factor limiting the rate of progress in plant
breeding has been low heritability.
 This occurs frequently in complex traits such as
yield and quality, which are controlled by a large
number of genes and are environmentally unstable.
 Plant breeders and physiologists are cooperating in
an effort to overcome this problem by breaking a
complex character down to components, and
ultimately to the actual molecules that mediate the
expression of the character.
 By selecting for the components of a character or
for its underlying molecular constitution, breeders
hope to reduce complexity and environmental
impact, allowing for faster progress.
 Areas currently being explored for application of
this approach are yield, quality, salt and drought
tolerance, water-logging tolerance chilling
tolerance, disease and insect resistance, and self-
incompatibility phenotypes (which are used in FI
hybrid production).
 For example, in processing tomatoes the major
component of yield is volume of finished product
(sauce, paste, ketchup) per unit of cultivated area.
 In breeding for harvest yield alone, much of the
progress observed results from increased water
content, which must, in turn, be eliminated by the
processor.
 The solution has been to maintain or increase the
economic yield of processing tomatoes
(solids)while decreasing water content.
 The alternative character used by breeders as a
selective criterion is sugar (soluble solids)
concentration, which is negatively correlated with
water content.
 By combining this character with additional
selection for gross yield, uniform maturity, fleshy
and thick skin it has been possible to make more
rapid progress in improving this crop than in the
past.
 Another approach to solving the problem of low
heritability, recently developed at the University of
California, is the use of electrophoretic variation as
an alternative selection criterion in tomatoes.
 One strength of cellular and molecular genetic
approaches is the possibility of performing selection
with chemicals or physical conditions on extremely
large populations of haploid individuals in a small
space and over a short time.
 Numerous schemes have been devised to select
desirable genetic variation with respect to nutrition,
stress tolerance, herbicide resistance, and disease
resistance.
 Until now, variant cells selected in vitro have only
rarely been translated into functionally equivalent
whole plants.
 Recent studies show that it may be possible to
circumvent some of the problems associated with
selection of cultured cells by substituting whole
plants.
 For example, chemicals have been used to screen
barley seedlings exhibiting increased nutritional
value.
 Other experiments have shown that pollen can be
responsive to selection in vivo, opening up the
possibility for screening large populations of haploid
individuals.
 Plant breeding has brought about drastic changes
in plant architecture or genetic makeup,
revolutionizing production and marketing methods.
 Recent examples include determinate growth habit
for mechanical harvesting, disease-resistant
cultivars, and hybrid varieties.
 New ideas are constantly emerging, such as
techniques to permit the selection of plants with
multiple tolerance genes (“horizontal” resistance) as
opposed to often unstable single gene (“vertical”)
resistance.
 Dwarf fruit trees are being developed that will
produce more with lower management and
harvesting costs.
 Although genetic engineering for crop improvement
holds great promise, plant breeding and other
aspects of whole plant genetics are also becoming
increasingly sophisticated.
 Advances in breeding technology guarantee that
plant breeders will continue to make valuable
contributions to agriculture.
 DESCRIPTION OF DIAGRAM
 To produce wheat-barley cross, embryo is excised
from immature barley endosperm (A) and is
replaced by hybrid embryo (B) produced by hand
pollination.
 The endosperm serves as nurse tissue for the
hybrid (C).
 About 70 percent of the embryo cultures may give
rise to plants (D), but of 270 plants obtained by this
process , only 20 were true wheat x barley hybrids
(E, center; spike at left is wheat; right, barley).
New concepts in maintenance of plant breeding promises and prospects
New concepts in maintenance of plant breeding promises and prospects
SOFTWARE FOR RAINFED NATU
TOBACCO GERMPLASM
 Among different tobacco types grown in India, rainfed
natu tobacco is an indigenous-type tobacco, mainly
used for domestic consumption; 81 rainfed natu tobacco
accessions are being maintained at the CTRI gene
bank.
 Based on the data available on various parameters of
these accessions, a knowledge-base system has been
developed with user-friendly menus using RDBMS
(relational database management system) technology
VB(visual basic).
 Net as Front-end and Oracle as Back-end application.
 This software helps store and retrieve data of tobacco
germplasm lines.
 The software consists of 40 parameters classified into
15 groups.
 The use of agro biodiversity for plant
improvement and the intellectual
property paradigm: institutional fit and
legal tools for mass selection,
conventional and molecular plant
breeding:-
Figure 1 Analyzing paradigmatic fit: expected
effects of the strong intellectual property
paradigm and partially open innovation in the
main fields of application of plant
improvement.
New concepts in maintenance of plant breeding promises and prospects
 Conclusion
 This paper has explored the effectiveness of the strong
intellectual property paradigm to create incentives for
innovation in the field of food and agricultural research,
relying on massive inputs of plant-genetic resources into
the research and development cycle.
 The main lesson is that there is not a single regime that
fits all contexts best at the same time.
 The strong paradigm has proven very effective in the
context of genetic engineering, but faces a systematic
paradigm breakdown when it extends its regulatory
scope over traditional mass selection operating for
example in exchange networks of farmers’ landraces,
where innovation often has a more collective,
community-related nature.
In these cases, a different form of intellectual-property
protection, based on partially-open innovation
systems, has proven more effective, even though
its very existence is threatened by the global
prevalence of the strong intellectual property
paradigm, pushed into forced illegality and faced
with significant disregard for the products of the
innovation process.
 Genetic engineering and mass selection with
naturally occurring landraces present two extreme
cases.
 Perhaps the most important part of modern
agricultural research is happening between the two
extremes of simple observation and deep molecular
introspection.
 Conventional breeding, characterized by controlled
hybridization, and based upon the methodically-
controlled crossing of specific varieties with other
plant-genetic material, implies the repeated input of
vast amounts of plant-genetic material, both from
the pool of already domesticated varieties and from
wild-plant genetic resources.
 Similarly, the use of molecular-biology-based gene
marker technologies or bio-informatics to empower
conventional breeding techniques is built around a
process of cumulative incremental innovation, where the
outputs of the research are used in turn as the direct
inputs for the next innovation cycle.
 For these intermediary categories of plant improvement
systems, both overly strong intellectual property rights
and the absence of well delineated intellectual-property
protection fail to provide incentives for investing in
follow-on innovations.
 In particular, in cases where access to plant-genetic
resources is a basic requirement to each new product
development cycle, overly strong intellectual property
rights might hamper or slow down the innovation
process due to increased transaction costs generated
by the need for complex licensing schemes for the use
of these resources.
 Further, due to the interdependence of developed
and developing countries for access to genetic
resources for food and agriculture – for example
situated in biodiversity hotspots in the South –
special attention is required to create investment in
genetic resources by countries that are situated far
from the innovation frontier and which are less well
placed to obtain and enforce strong intellectual
property rights.
 Intellectual property rights legislation ought in this
regard be recalibrated so as to incorporate farmers’
needs and rights, all the while constructing a well-
suited reward regime to maintain adequate levels of
genetic diversity that are not only important for
farmers’ livelihoods or for environmental
conservation policies, but also for the future of
agricultural research and development efforts
worldwide.
 As can be seen from the analysis in this article,
neither strong intellectual property rights, nor their
complete absence have proven an optimal fit from a
legal and economic point view for the two
intermediary categories of cumulative incremental
innovation in agriculture research.
 Finally, this conclusion leads us to recognize the
role played by other arguments, beyond the legal
analysis of optimal fit, when making regulatory
decisions for intermediary categories of cumulative
incremental innovation.
 Broader collective decisions on investments in the
organizational and institutional infrastructures in
scientific research will play a role in the choice of
the most adequate legal framework for agro
biodiversity-based innovation in a given society.
For instance, broader societal choices to be made for
tackling the challenges posed by climate change for
world-food security, may lead to decisions to
increase investment in transgenic technologies
developed either by private sector entities or by the
public sector if granted sufficient funds to do so.
It could also lead to increased investment in global
collaboration between both public and private
research institutions for experimental breeding, as a
way to tackle these challenges.
BREEDING STRATEGIES FOR THE
IMPROVEMENT OF PULSE CROPS
 Select the plant having high biological yield coupled with
high harvest index.
 Search variety having continuous photosynthesizing
activity with more branches, and pods leads to
productive genotypes with high harvest index.
 More vegetative growth is limiting factor for poor grain
yield.
 PLANT TYPE CONCEPT:
 Improved plant type can increase the harvest efficiency.
 Improvement of plant type by genetic restructuring in the
direction of high harvest index.
 Improvement is done by genetic manipulation.
 Restructuring of the canopy so that greater part of it
receives sufficient energy , may results in high yield.
 CHARACTERS ASSOCIATION: Yield is associated with
various characters like no. of branches, no. of pods,
seeds per pod, pod per plant and seed size. These
characters are related to each other. Optimum
improvement of these characters helpful in increasing
yield.
 TYPE OF GENE ACTION:
 Both additive and non additive gene effects recorded.
 Yield and its characters associated with dominance
gene effect.
 Contribution of complementary gene were also
substantial.
 Complementary epistatis important for yield,
podding and seed number can be exploited for crop
improvement.
 Multiple crosses followed by intermating among
desirable segregants for improvement of crop yield.
 The progress in breeding of self pollinated crops is
relatively slow.
 The conventional method is pedigree method but to
maintain the variability among varieties bulk method
is used.
SELECTION OF PROSPECTIVE PARENT AMONG
INDIAN AND EXOTIC SESAME
 Marker assisted selection (MAS) include:
 Isolation of genomic DNA
 RAPD analysis
 SCAR marker development
 SSR analysis
 Band scoring and analysis
New concepts in maintenance of plant breeding promises and prospects

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New concepts in maintenance of plant breeding promises and prospects

  • 2. NEW CONCEPTS IN MAINTENANCE OF PLANT BREEDING PROMISES AND PROSPECTS SUBMITTED TO Dr. M.REDDI SEKHAR PRESENTED BY ZUBY GOHAR ANSARI TAM/14/26
  • 3.  NEW CONCEPTS IN WHOLE PLANT GENETICS  In the excitement surrounding accomplishments in the manipulation of tissues, cells, and genes over the past ten years, one might get the impression that plant breeding research at the whole plant and population levels has been fixed away and forgotten.  On the contrary, several exciting developments in breeding technology have surfaced during this same period.  One premise upon which the need for cellular and molecular approaches has been based is that plant breeders have exhausted existing pools of genetic variability, and progress has consequently diminished.
  • 4.  There is little evidence to substantiate the existence of such limitations for most crops. Moreover, it is premature to conclude that sexually accessible variability has been depleted when the existing collections are rarely, if ever, comprehensive.  New variability, for example, in soybean and rice introductions (among others) from the People’s Republic of China will more likely sustain performance increases in the foreseeable future than will new cellular/molecular approaches.
  • 5.  The idea of expanding the pool of accessible genes by overcoming natural isolating mechanisms between species remains a possibility; hybrids between species can be synthesized and chromosomally altered to produce dramatic economic gains, as with strawberries; it is now possible to sexually engineer wheat lines containing desired genes from related species simply by adding or substituting whole chromosomes.
  • 6.  In other research, crosses between the cultivated tomato and a salt-tolerant wild species have produced breeding lines with some salt tolerance.  Preliminary observations suggest that cold tolerance is another possibility.  To date, 14 genes for disease resistance have been transferred from wild species into cultivated tomato varieties.  Cell culture of interspecific hybrids has been successfully applied to facilitate the flow of genes among species related to tomato, barley, and cabbage.  Altogether, exploitation of genes from wild germplasm sources has barely begun.
  • 8.  A major factor limiting the rate of progress in plant breeding has been low heritability.  This occurs frequently in complex traits such as yield and quality, which are controlled by a large number of genes and are environmentally unstable.  Plant breeders and physiologists are cooperating in an effort to overcome this problem by breaking a complex character down to components, and ultimately to the actual molecules that mediate the expression of the character.  By selecting for the components of a character or for its underlying molecular constitution, breeders hope to reduce complexity and environmental impact, allowing for faster progress.
  • 9.  Areas currently being explored for application of this approach are yield, quality, salt and drought tolerance, water-logging tolerance chilling tolerance, disease and insect resistance, and self- incompatibility phenotypes (which are used in FI hybrid production).  For example, in processing tomatoes the major component of yield is volume of finished product (sauce, paste, ketchup) per unit of cultivated area.  In breeding for harvest yield alone, much of the progress observed results from increased water content, which must, in turn, be eliminated by the processor.
  • 10.  The solution has been to maintain or increase the economic yield of processing tomatoes (solids)while decreasing water content.  The alternative character used by breeders as a selective criterion is sugar (soluble solids) concentration, which is negatively correlated with water content.  By combining this character with additional selection for gross yield, uniform maturity, fleshy and thick skin it has been possible to make more rapid progress in improving this crop than in the past.
  • 11.  Another approach to solving the problem of low heritability, recently developed at the University of California, is the use of electrophoretic variation as an alternative selection criterion in tomatoes.  One strength of cellular and molecular genetic approaches is the possibility of performing selection with chemicals or physical conditions on extremely large populations of haploid individuals in a small space and over a short time.  Numerous schemes have been devised to select desirable genetic variation with respect to nutrition, stress tolerance, herbicide resistance, and disease resistance.
  • 12.  Until now, variant cells selected in vitro have only rarely been translated into functionally equivalent whole plants.  Recent studies show that it may be possible to circumvent some of the problems associated with selection of cultured cells by substituting whole plants.  For example, chemicals have been used to screen barley seedlings exhibiting increased nutritional value.  Other experiments have shown that pollen can be responsive to selection in vivo, opening up the possibility for screening large populations of haploid individuals.
  • 13.  Plant breeding has brought about drastic changes in plant architecture or genetic makeup, revolutionizing production and marketing methods.  Recent examples include determinate growth habit for mechanical harvesting, disease-resistant cultivars, and hybrid varieties.  New ideas are constantly emerging, such as techniques to permit the selection of plants with multiple tolerance genes (“horizontal” resistance) as opposed to often unstable single gene (“vertical”) resistance.  Dwarf fruit trees are being developed that will produce more with lower management and harvesting costs.
  • 14.  Although genetic engineering for crop improvement holds great promise, plant breeding and other aspects of whole plant genetics are also becoming increasingly sophisticated.  Advances in breeding technology guarantee that plant breeders will continue to make valuable contributions to agriculture.
  • 15.  DESCRIPTION OF DIAGRAM  To produce wheat-barley cross, embryo is excised from immature barley endosperm (A) and is replaced by hybrid embryo (B) produced by hand pollination.  The endosperm serves as nurse tissue for the hybrid (C).  About 70 percent of the embryo cultures may give rise to plants (D), but of 270 plants obtained by this process , only 20 were true wheat x barley hybrids (E, center; spike at left is wheat; right, barley).
  • 18. SOFTWARE FOR RAINFED NATU TOBACCO GERMPLASM  Among different tobacco types grown in India, rainfed natu tobacco is an indigenous-type tobacco, mainly used for domestic consumption; 81 rainfed natu tobacco accessions are being maintained at the CTRI gene bank.  Based on the data available on various parameters of these accessions, a knowledge-base system has been developed with user-friendly menus using RDBMS (relational database management system) technology VB(visual basic).  Net as Front-end and Oracle as Back-end application.  This software helps store and retrieve data of tobacco germplasm lines.  The software consists of 40 parameters classified into 15 groups.
  • 19.  The use of agro biodiversity for plant improvement and the intellectual property paradigm: institutional fit and legal tools for mass selection, conventional and molecular plant breeding:- Figure 1 Analyzing paradigmatic fit: expected effects of the strong intellectual property paradigm and partially open innovation in the main fields of application of plant improvement.
  • 21.  Conclusion  This paper has explored the effectiveness of the strong intellectual property paradigm to create incentives for innovation in the field of food and agricultural research, relying on massive inputs of plant-genetic resources into the research and development cycle.  The main lesson is that there is not a single regime that fits all contexts best at the same time.  The strong paradigm has proven very effective in the context of genetic engineering, but faces a systematic paradigm breakdown when it extends its regulatory scope over traditional mass selection operating for example in exchange networks of farmers’ landraces, where innovation often has a more collective, community-related nature.
  • 22. In these cases, a different form of intellectual-property protection, based on partially-open innovation systems, has proven more effective, even though its very existence is threatened by the global prevalence of the strong intellectual property paradigm, pushed into forced illegality and faced with significant disregard for the products of the innovation process.  Genetic engineering and mass selection with naturally occurring landraces present two extreme cases.
  • 23.  Perhaps the most important part of modern agricultural research is happening between the two extremes of simple observation and deep molecular introspection.  Conventional breeding, characterized by controlled hybridization, and based upon the methodically- controlled crossing of specific varieties with other plant-genetic material, implies the repeated input of vast amounts of plant-genetic material, both from the pool of already domesticated varieties and from wild-plant genetic resources.
  • 24.  Similarly, the use of molecular-biology-based gene marker technologies or bio-informatics to empower conventional breeding techniques is built around a process of cumulative incremental innovation, where the outputs of the research are used in turn as the direct inputs for the next innovation cycle.  For these intermediary categories of plant improvement systems, both overly strong intellectual property rights and the absence of well delineated intellectual-property protection fail to provide incentives for investing in follow-on innovations.  In particular, in cases where access to plant-genetic resources is a basic requirement to each new product development cycle, overly strong intellectual property rights might hamper or slow down the innovation process due to increased transaction costs generated by the need for complex licensing schemes for the use of these resources.
  • 25.  Further, due to the interdependence of developed and developing countries for access to genetic resources for food and agriculture – for example situated in biodiversity hotspots in the South – special attention is required to create investment in genetic resources by countries that are situated far from the innovation frontier and which are less well placed to obtain and enforce strong intellectual property rights.  Intellectual property rights legislation ought in this regard be recalibrated so as to incorporate farmers’ needs and rights, all the while constructing a well- suited reward regime to maintain adequate levels of genetic diversity that are not only important for farmers’ livelihoods or for environmental conservation policies, but also for the future of agricultural research and development efforts worldwide.
  • 26.  As can be seen from the analysis in this article, neither strong intellectual property rights, nor their complete absence have proven an optimal fit from a legal and economic point view for the two intermediary categories of cumulative incremental innovation in agriculture research.  Finally, this conclusion leads us to recognize the role played by other arguments, beyond the legal analysis of optimal fit, when making regulatory decisions for intermediary categories of cumulative incremental innovation.  Broader collective decisions on investments in the organizational and institutional infrastructures in scientific research will play a role in the choice of the most adequate legal framework for agro biodiversity-based innovation in a given society.
  • 27. For instance, broader societal choices to be made for tackling the challenges posed by climate change for world-food security, may lead to decisions to increase investment in transgenic technologies developed either by private sector entities or by the public sector if granted sufficient funds to do so. It could also lead to increased investment in global collaboration between both public and private research institutions for experimental breeding, as a way to tackle these challenges.
  • 28. BREEDING STRATEGIES FOR THE IMPROVEMENT OF PULSE CROPS  Select the plant having high biological yield coupled with high harvest index.  Search variety having continuous photosynthesizing activity with more branches, and pods leads to productive genotypes with high harvest index.  More vegetative growth is limiting factor for poor grain yield.  PLANT TYPE CONCEPT:  Improved plant type can increase the harvest efficiency.  Improvement of plant type by genetic restructuring in the direction of high harvest index.
  • 29.  Improvement is done by genetic manipulation.  Restructuring of the canopy so that greater part of it receives sufficient energy , may results in high yield.  CHARACTERS ASSOCIATION: Yield is associated with various characters like no. of branches, no. of pods, seeds per pod, pod per plant and seed size. These characters are related to each other. Optimum improvement of these characters helpful in increasing yield.  TYPE OF GENE ACTION:  Both additive and non additive gene effects recorded.  Yield and its characters associated with dominance gene effect.  Contribution of complementary gene were also substantial.
  • 30.  Complementary epistatis important for yield, podding and seed number can be exploited for crop improvement.  Multiple crosses followed by intermating among desirable segregants for improvement of crop yield.  The progress in breeding of self pollinated crops is relatively slow.  The conventional method is pedigree method but to maintain the variability among varieties bulk method is used.
  • 31. SELECTION OF PROSPECTIVE PARENT AMONG INDIAN AND EXOTIC SESAME  Marker assisted selection (MAS) include:  Isolation of genomic DNA  RAPD analysis  SCAR marker development  SSR analysis  Band scoring and analysis