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Evolution
                    by
             Natural Selection




AP Biology                       2006-2007
DOCTRINE
AP Biology
           TINTORETTO   The Creation of the Animals 1550
But the Fossil record…




             OBSERVATION




AP Biology
mya
 Quaternary




                                                                                                           Molluscs
                      Anaerobic Bacteria




                                                                                                                                               Jawless Fish

                                                                                                                                                              Teleost Fish
                                                                                   Multicellular Animals




                                                                                                                                   Chordates




                                                                                                                                                                                                                                             Land Plants
                                           Photosynthetic Bacteria

                                                                     Green Algae




                                                                                                                                                                                                                                                           Seed Plants
                                                                                                                      Arthropods




                                                                                                                                                                             Amphibians




                                                                                                                                                                                                                           Mammals

                                                                                                                                                                                                                                     Birds




                                                                                                                                                                                                                                                                         Flowering Plants
                                                                                                                                                                                          Insects

                                                                                                                                                                                                    Reptiles
              1.5
 Tertiary
                 63
 Cretaceous




                                                                                                                                                                                                               Dinosaurs
              135
 Jurassic
              180
 Triassic
              225
 Permian
              280
 Carboniferous
              350
 Devonian
              400
 Silurian
              430
 Ordovician
              500
 Cambrian
              570
 Ediacaran
              700
 Precambrian,
 Proterozoic,
       &
 Archarozoic
              4500

Life’s Natural History is a record of Successions & Extinctions
  AP Biology
LaMarck
    Organisms adapted to
      their environments by
      acquiring traits
            change in their life time
               Disuse
                organisms lost parts because they did not use them
                — like the missing eyes & digestive system of the
                tapeworm
               Perfection with Use & Need
                the constant use of an organ leads that organ to
                increase in size — like the muscles of a blacksmith
                or the large ears of a night-flying bat
            transmit acquired characteristics to next
             generation
AP Biology
Charles Darwin
    1809-1882
    British naturalist
    Proposed the idea
      of evolution by
      natural selection
     Collected clear
      evidence to
      support his ideas


AP Biology
Voyage of the HMS Beagle
    Invited to travel around the world
        1831-1836 (22 years old!)
        makes many observations of nature

              main mission of the Beagle was to chart
              South American coastline



  Robert Fitzroy




AP Biology
Voyage of the HMS Beagle
     Stopped in Galapagos Islands
            500 miles off coast of Ecuador




AP Biology
Succession of types
    Armadillos are native to the
    Americas, with most species
    found in South America.




                               Why should extinct
                              armadillo-like species
                              & living armadillos be
                               found on the same
                                    continent?
Glyptodont fossils are also
unique to South America.
 AP Biology
Mylodon (left) Giant
                                        ground sloth (extinct)




             Modern sloth (right)


                             “This wonderful relationship
                           in the same continent between
                    the dead and the living will…throw more light
                  on the appearance of organic beings on our earth,
                          and their disappearance from it,
                           than any other class of facts.”
AP Biology
Unique species




AP Biology
Darwin found… birds
     Collected many
     different birds on the
     Galapagos Islands.


                           Finch?       Sparrow?
     Thought he found
     very different kinds…




AP Biology                Woodpecker?   Warbler?
But Darwin found… a lot of finches
Darwin was amazed to
find out:
All 14 species of birds
were finches…
But there is only one
species of finch on the   LargeFinch?
                                Ground
                              Finch?         Sparrow?
                                            Small Ground
                                             Sparrow?
mainland!                     Finch            Finch


                   How did
                 one species
              of finches become
              so many different
                species now?

AP Biology                 Woodpecker?
                           Warbler Finch
                           Woodpecker?        Warbler?
                                           Veg. Tree Finch
                                              Warbler?
Tree Thinking
Descendant
 species




   Ancestral       Large-seed eater?
                     Large Ground      Small-seed eater?
                                         Small Ground
    species              Finch               Finch




AP Biology            Warbler?
                      Warbler Finch     Leaf-browser?
                                        Veg. Tree Finch
Correlation of species to food source




     Seed        Flower            Insect
    eaters       eaters            eaters

                                      Rapid speciation:
                               new species filling new niches,
                                   because they inherited
                                  successful adaptations.




Adaptive radiation
 AP Biology
Darwin’s finches
    Differences in beaks
          associated with eating different foods
          survival & reproduction of beneficial

           adaptations to foods available on islands
                                 Warbler finch       Cactus finch

             Woodpecker finch                                        Sharp-beaked finch

  Small insectivorous                           er                            Small ground
            tree finch                     Warbl                              finch
                                            finch



                                                             Gr
                                     es




           Large                                 Cactus



                                                                ou
                                   ch




   insectivorous                                                                 Medium



                                                                  nd
                                fin




                                                  eater
       tree finch                                                                ground finch


                                                                    fin
                             ee




                                     Insect eaters
                          Tr




                                                                       ch
                                                     Seed eaters

                                                                         es
    Vegetarian
                                  Bud eater                                       Large
     tree finch
AP Biology                                                                        ground finch
Darwin’s finches
   Darwin’s conclusions
          small populations of original South American
           finches landed on islands
              variation in beaks enabled individuals to gather
              food successfully in the different environments
          over many generations, the populations of
           finches changed anatomically & behaviorally
              accumulation of advantageous traits in population
              emergence of different species



AP Biology
Seeing this gradation &
                        diversity of structure in
             one small, intimately related group of birds,
                      one might really fancy that
                   from an original paucity of birds
                           in this archipelago,
                     one species has been taken &
                      modified for different ends.




AP Biology
Darwin’s finches
     Differences in
       beaks allowed
       some finches to…
         successfully
          compete
         successfully feed

         successfully

          reproduce
              pass successful
              traits onto their
              offspring
AP Biology
More observations…   Correlation of species
                             to food source




                          Whoa,
                       Turtles, too!



 AP Biology
Many islands also show
                  distinct local variations in
                  tortoise morphology…


              …perhaps these are
             the first steps in the
             splitting of one species
                   into several?
AP Biology
Artificial selection




This is not just a
process of the
past…
                                        It is all
                                        around
AP Biology                              us today
Selective
   breeding
   the raw genetic
   material (variation)
   is hidden there




AP Biology
Selective breeding
      Hidden variation
      can be exposed
      through selection!




AP Biology
In historical context
      Other people’s ideas paved the
       path for Darwin’s thinking


                                       competition:
                                   struggle for survival
                                    population growth
                                   exceeds food supply




 land masses change over
    immeasurable time
AP Biology
A Reluctant Revolutionary
    Returned to England in 1836
          wrote papers describing his collections
           & observations
          long treatise on barnacles

          draft of his theory of

           species formation in 1844
              instructed his wife to
               publish this essay upon
               his death
              reluctant to publish but
               didn’t want ideas to die
AP Biology     with him
And then came the letter….
   Then, in 1858, Darwin received a letter
   that changed everything…


      Alfred Russel Wallace
      a young naturalist working
      in the East Indies, had
      written a short paper with a
      new idea. He asked Darwin
      to evaluate his ideas and
      pass it along for publication.


AP Biology
The time was ripe for the idea!




                     To Lyell—
                         Your words
                      have come true
                     with a vengeance…
                I never saw a more striking
             coincidence…so all my originality,
                whatever it may amount to,
                      will be smashed.


AP Biology
Voyage: 1831-1836

    November 24, 1859, Darwin published
“On the Origin of Species by Means of Natural Selection”




 AP Biology
Essence of Darwin’s ideas
    Natural selection
        variation exists in populations
        over-production of offspring

               more offspring than the environment can support
            competition
               for food, mates, nesting sites, escape predators
            differential survival
               successful traits = adaptations
            differential reproduction
               adaptations become more
AP Biology
               common in population
LaMarckian vs. Darwinian view
    LaMarck
            in reaching higher
             vegetation giraffes
             stretch their necks &
             transmits the acquired
             longer neck to offspring
     Darwin



                 
            giraffes born with longer
             necks survive better &
             leave more offspring who
             inherit their long necks
AP Biology
Stick your neck out…
           Ask Questions!
AP Biology
Ghosts of Lectures Past
                    (storage)




AP Biology                             2009-2010
Life has changed over time
       & in turn has changed the Earth




                              Living creatures have
                          changed Earth’s environment,
                            making other life possible




AP Biology
Evolution as Change Over Time



                        idea accepted
             Evolution!
                        before Darwin
                                        Evolution!


                Evolution!


                       Evolution!
AP Biology
Galapagos
    Recent volcanic origin most
    of animal species on the
    Galápagos live nowhere else
    in world, but they resemble
    species living on South
    American mainland.




AP Biology                        500 miles west of mainland
Coherent explanation of observations




                "Nothing in biology
                makes sense except in
                the light of evolution."
                     -- Theodosius Dobzhansky
                                         March 1973
                      Geneticist, Columbia University
                                         (1900-1975)
AP Biology                                   2006-2007
Essence of Darwin’s ideas
              (1) Variation exists in natural populations
              (2) Many more offspring are born each season
                  than can possibly survive to maturity
              (3) As a result, there is a struggle for existence
                  - competition
               (4) Characteristics beneficial in the struggle
                   for existence will tend to become more
                   common in the population, changing the
                   average characteristics of the population
                    - adaptations
(5) Over long periods of time, and given a steady input of
    new variation into a population, these processes lead to
AP Biologyemergence of new species
    the
Stick your neck out…
                Ask Questions!




AP Biology
The Birds…
     Galápagos birds
            22 of the 29 species of
             birds on the Galapagos
             are endemic
               found only on these islands
            collected specimens of all
     One particular group…
            at first, he paid little
             attention to a series of
             small birds
            some were woodpecker-
             like, some warbler-like, &
             some finch-like

AP Biology
Darwin’s finches
    Darwin was amazed
      to find out they were
      all finches
            14 species
            but only one
             species on SouthLarge-seed eater?
                                   Finch?               Small-seed eater?
                                                            Sparrow?
             American mainland
               500 miles away
            all the birds had to
             originally come             How did
             from mainland             one species
                                    of finches become
             species                so many different
                                        ones now?
AP Biology                          Warbler?
                                     Warbler?            Leaf-browser?
                                                             Wren?

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02 ch22evolutiondarwin2009

  • 1. Evolution by Natural Selection AP Biology 2006-2007
  • 2. DOCTRINE AP Biology TINTORETTO The Creation of the Animals 1550
  • 3. But the Fossil record… OBSERVATION AP Biology
  • 4. mya Quaternary Molluscs Anaerobic Bacteria Jawless Fish Teleost Fish Multicellular Animals Chordates Land Plants Photosynthetic Bacteria Green Algae Seed Plants Arthropods Amphibians Mammals Birds Flowering Plants Insects Reptiles 1.5 Tertiary 63 Cretaceous Dinosaurs 135 Jurassic 180 Triassic 225 Permian 280 Carboniferous 350 Devonian 400 Silurian 430 Ordovician 500 Cambrian 570 Ediacaran 700 Precambrian, Proterozoic, & Archarozoic 4500 Life’s Natural History is a record of Successions & Extinctions AP Biology
  • 5. LaMarck  Organisms adapted to their environments by acquiring traits  change in their life time  Disuse organisms lost parts because they did not use them — like the missing eyes & digestive system of the tapeworm  Perfection with Use & Need the constant use of an organ leads that organ to increase in size — like the muscles of a blacksmith or the large ears of a night-flying bat  transmit acquired characteristics to next generation AP Biology
  • 6. Charles Darwin  1809-1882  British naturalist  Proposed the idea of evolution by natural selection  Collected clear evidence to support his ideas AP Biology
  • 7. Voyage of the HMS Beagle  Invited to travel around the world  1831-1836 (22 years old!)  makes many observations of nature  main mission of the Beagle was to chart South American coastline Robert Fitzroy AP Biology
  • 8. Voyage of the HMS Beagle  Stopped in Galapagos Islands  500 miles off coast of Ecuador AP Biology
  • 9. Succession of types Armadillos are native to the Americas, with most species found in South America. Why should extinct armadillo-like species & living armadillos be found on the same continent? Glyptodont fossils are also unique to South America. AP Biology
  • 10. Mylodon (left) Giant ground sloth (extinct) Modern sloth (right) “This wonderful relationship in the same continent between the dead and the living will…throw more light on the appearance of organic beings on our earth, and their disappearance from it, than any other class of facts.” AP Biology
  • 12. Darwin found… birds Collected many different birds on the Galapagos Islands. Finch? Sparrow? Thought he found very different kinds… AP Biology Woodpecker? Warbler?
  • 13. But Darwin found… a lot of finches Darwin was amazed to find out: All 14 species of birds were finches… But there is only one species of finch on the LargeFinch? Ground Finch? Sparrow? Small Ground Sparrow? mainland! Finch Finch How did one species of finches become so many different species now? AP Biology Woodpecker? Warbler Finch Woodpecker? Warbler? Veg. Tree Finch Warbler?
  • 14. Tree Thinking Descendant species Ancestral Large-seed eater? Large Ground Small-seed eater? Small Ground species Finch Finch AP Biology Warbler? Warbler Finch Leaf-browser? Veg. Tree Finch
  • 15. Correlation of species to food source Seed Flower Insect eaters eaters eaters Rapid speciation: new species filling new niches, because they inherited successful adaptations. Adaptive radiation AP Biology
  • 16. Darwin’s finches  Differences in beaks  associated with eating different foods  survival & reproduction of beneficial adaptations to foods available on islands Warbler finch Cactus finch Woodpecker finch Sharp-beaked finch Small insectivorous er Small ground tree finch Warbl finch finch Gr es Large Cactus ou ch insectivorous Medium nd fin eater tree finch ground finch fin ee Insect eaters Tr ch Seed eaters es Vegetarian Bud eater Large tree finch AP Biology ground finch
  • 17. Darwin’s finches  Darwin’s conclusions  small populations of original South American finches landed on islands  variation in beaks enabled individuals to gather food successfully in the different environments  over many generations, the populations of finches changed anatomically & behaviorally  accumulation of advantageous traits in population  emergence of different species AP Biology
  • 18. Seeing this gradation & diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species has been taken & modified for different ends. AP Biology
  • 19. Darwin’s finches  Differences in beaks allowed some finches to…  successfully compete  successfully feed  successfully reproduce  pass successful traits onto their offspring AP Biology
  • 20. More observations… Correlation of species to food source Whoa, Turtles, too! AP Biology
  • 21. Many islands also show distinct local variations in tortoise morphology… …perhaps these are the first steps in the splitting of one species into several? AP Biology
  • 22. Artificial selection This is not just a process of the past… It is all around AP Biology us today
  • 23. Selective breeding the raw genetic material (variation) is hidden there AP Biology
  • 24. Selective breeding Hidden variation can be exposed through selection! AP Biology
  • 25. In historical context  Other people’s ideas paved the path for Darwin’s thinking competition: struggle for survival population growth exceeds food supply land masses change over immeasurable time AP Biology
  • 26. A Reluctant Revolutionary  Returned to England in 1836  wrote papers describing his collections & observations  long treatise on barnacles  draft of his theory of species formation in 1844  instructed his wife to publish this essay upon his death  reluctant to publish but didn’t want ideas to die AP Biology with him
  • 27. And then came the letter…. Then, in 1858, Darwin received a letter that changed everything… Alfred Russel Wallace a young naturalist working in the East Indies, had written a short paper with a new idea. He asked Darwin to evaluate his ideas and pass it along for publication. AP Biology
  • 28. The time was ripe for the idea! To Lyell— Your words have come true with a vengeance… I never saw a more striking coincidence…so all my originality, whatever it may amount to, will be smashed. AP Biology
  • 29. Voyage: 1831-1836 November 24, 1859, Darwin published “On the Origin of Species by Means of Natural Selection” AP Biology
  • 30. Essence of Darwin’s ideas  Natural selection  variation exists in populations  over-production of offspring  more offspring than the environment can support  competition  for food, mates, nesting sites, escape predators  differential survival  successful traits = adaptations  differential reproduction  adaptations become more AP Biology common in population
  • 31. LaMarckian vs. Darwinian view  LaMarck  in reaching higher vegetation giraffes stretch their necks & transmits the acquired longer neck to offspring  Darwin   giraffes born with longer necks survive better & leave more offspring who inherit their long necks AP Biology
  • 32. Stick your neck out… Ask Questions! AP Biology
  • 33. Ghosts of Lectures Past (storage) AP Biology 2009-2010
  • 34. Life has changed over time & in turn has changed the Earth Living creatures have changed Earth’s environment, making other life possible AP Biology
  • 35. Evolution as Change Over Time idea accepted Evolution! before Darwin Evolution! Evolution! Evolution! AP Biology
  • 36. Galapagos Recent volcanic origin most of animal species on the Galápagos live nowhere else in world, but they resemble species living on South American mainland. AP Biology 500 miles west of mainland
  • 37. Coherent explanation of observations "Nothing in biology makes sense except in the light of evolution." -- Theodosius Dobzhansky March 1973 Geneticist, Columbia University (1900-1975) AP Biology 2006-2007
  • 38. Essence of Darwin’s ideas (1) Variation exists in natural populations (2) Many more offspring are born each season than can possibly survive to maturity (3) As a result, there is a struggle for existence - competition (4) Characteristics beneficial in the struggle for existence will tend to become more common in the population, changing the average characteristics of the population - adaptations (5) Over long periods of time, and given a steady input of new variation into a population, these processes lead to AP Biologyemergence of new species the
  • 39. Stick your neck out… Ask Questions! AP Biology
  • 40. The Birds…  Galápagos birds  22 of the 29 species of birds on the Galapagos are endemic  found only on these islands  collected specimens of all  One particular group…  at first, he paid little attention to a series of small birds  some were woodpecker- like, some warbler-like, & some finch-like AP Biology
  • 41. Darwin’s finches  Darwin was amazed to find out they were all finches  14 species  but only one species on SouthLarge-seed eater? Finch? Small-seed eater? Sparrow? American mainland  500 miles away  all the birds had to originally come How did from mainland one species of finches become species so many different ones now? AP Biology Warbler? Warbler? Leaf-browser? Wren?

Hinweis der Redaktion

  1. Lamarck noted how well-adapted organisms were to their environments, and believed that fossils could be understood as less perfect forms which had perished in the struggle for increasing perfection. He explained adaptation as a result of change caused by environmental pressures.
  2. What did Darwin say? What evidence supports Evolution by Natural Selection? What impact did Evolution have on biology?
  3. After graduation Darwin was recommended to be the conversation companion to Captain Robert FitzRoy, preparing the survey ship Beagle for a voyage around the world. FitzRoy chose Darwin because of his education, his similar social class, and similar age as the captain. Darwin noted that the plants and animals of South America were very distinct from those of Europe
  4. The origin of the fauna of the Galapagos, 900 km west of the South American coast, especially puzzled Darwin. On further study after his voyage, Darwin noted that while most of the animal species on the Galapagos lived nowhere else, they resembled species living on the South American mainland. It seemed that the islands had been colonized by plants and animals from the mainland that had then diversified on the different islands
  5. Show Campbell videos!!!
  6. Darwin noted that the plants and animals of South America were very distinct from those of Europe. Organisms from temperate regions of South America were more similar to those from the tropics of South America than to those from temperate regions of Europe. Further, South American fossils more closely resembled modern species from that continent than those from Europe.
  7. Theodosius Dobzhansky: Integrating Genetics and Evolution Theodosius Dobzhansky, a Russian geneticist who moved to the United States, provided laboratory evidence for natural selection and variation where previously there had been only field observation. Dobzhansky's work with Drosophila, or fruit flies, provided new evidence that supported Darwin's theory that natural selection, acting on genetic variation in populations, is a driving force in evolution.
  8. Darwin noted that the plants and animals of South America were very distinct from those of Europe. Organisms from temperate regions of South America were more similar to those from the tropics of South America than to those from temperate regions of Europe. Further, South American fossils more closely resembled modern species from that continent than those from Europe.