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Evolution

                         Popular Belief
• Prior to the studies of Charles Darwin, the most
  widespread belief was that all known species were
  created at the same time and remained unchanged
  throughout history.
• Some scientists at the time believed that features an
  individual acquired during a lifetime could be passed
  onto its offspring, and the species could gradually
  change to fit an environment better.

       A scientific theory is NOT just a belief, it is a well-supported tested
       explanation.

 2.1
Evolution

                  Descent with Modification




    Evolution means change over time
Evolution can also be defined more narrowly as a change in the genetic
composition of a population from generation to generation.
Fossils
• A fossil is the preserved   • Fossils are found in
  remains or imprints of        sedimentary rock
  ancient organisms.            formed when sand and
                                silt settle to the bottom
• Someone who studies           of water.
  fossils is called a
  paleontologist.
Types of radioactive dating
                                • Carbon dating: 2 types
• Potassium-Argon dating is
  used for rocks that are       • 14C:12C
  suspected billions of years      – Radioactive 14C
  old.                             – Non-radioactive 12C, most
• In older rocks the ratio of        common
  Argon to 40K will be higher
  than in younger rocks.
Evidences for Evolution                      Fossil Record
                                                 claws
                                                          wing-like
                                                          forelimbs




                               teeth
                                                                           thin
                                                                           ribs

                                   long tail


                                                                    feathers

                                               Reptile-like features
Replica of Archaeopteryx fossil;
     half bird half reptile                    Bird-like features
Evidence from comparative Anatomy

                     Vertebrate Limbs

The basic pattern of the vertebrate limb is represented by the
             human arm skeleton shown below

ball and                                     group of
                     hinge joint             small
socket joint
                                             bones




   single bone                two
                              bones             five jointed
                                                digits
19
The forelimbs of the following vertebrates show the basic
pattern of limb bones with modifications which are adapted
to their methods of locomotion.

The basic pattern suggests they have evolved from a common
                           ancestor




     Lizard                             Bird




    Dolphin                            Bat
DNA evidence

  Another important line of evidence for evolution comes
  from DNA analysis


Organisms which have much of their DNA in common must be
closely related, i.e. they have split from a common ancestor
comparatively recently (in geological terms)


For example, humans and chimpanzees have 99% of
their DNA in common which suggests a close
relationship and relatively ‘recent’ divergence from a
common ancestor
Evolutionary Theory

Evolution is the process in which significant changes
in the inheritable traits (genetic makeup) of a species
occur over time.

The validity and value of evolutionary biology are
based on continual analysis and interpretation of
accumulating science data.


    In words of Charles Darwin evolution is Descent with
                        modification
Theories of Organic Evolution


 Historically three theories
1. Lamarck’s Theory of Evolution, 2. Darwin’s Theory of
   Evolution, 3. De Vries Mutation Theory of Evolution.

 The views of most modern biologists combine the second
  and third

 No evidence of the first theory, that of Lamarck’s Use
  and disuse theory.
Jean Baptiste Lamarck, 1744-1829


               • French geologist and naturalist

               • published Philosophie
                 Zoologique in 1809

               • Speculated about mechanisms
                 of biological evolution, offered
                 no evidence
In 1809 Jean-Baptiste Lamarke hypothesized that “simple”
organisms emerge spontaneously and evolve to greater
“complexity”.

                Principles to his theory

      1. Organisms constantly strive to improve
         themselves.
      2. Most used body structures develop and
         unused waste away. (Use and Disuse)
      3. A structure is modified by use or disuse, the
         modification is inherited to the offspring.
         (Inheritance of acquired characteristics).
• Structural variations are due to
  functional needs – “to use or
  disuse”
• Use of a structure increases its size.
• Failure to use it decreases its size
• These variations (now referred to as
  “acquired characteristics” are
  inherited.
• There is no supporting evidence in
  modern biology to support
  Lamarck’s idea
Disproving Lamarck
• Characteristics acquired during the lifetime of a parent are not
  passed onto the offspring

• An athlete who develops a large muscle mass through training
  does not have children who already possess this large muscle
  mass

• Ernst Haeckel
  In an attempt to disprove Lamarckism he is said to have cut off
  the tails of mice for several generations

• The babies born from this line of tailless mice still grew tails
  as long as their ancestors

• This was not exactly a fair test as the mice had not stopped
  using their tails in an attempt to adapt to their environment

•
Charles Darwin: The Theory of Natural Selection

• His theory was effectively
  presented to the world in
  1859 when his book “The
  Origin of Species” was
  published.

• Darwin’s Theory rests on 5
  principles:
5 principles of Darwin’s Theory

1. Overproduction
   - All species have a tendency and the potential to
   increase at a geometric rate.
All organisms produce more organisms but only a few
   survive.
2. Competition
   - The conditions supporting life are limited.
      For eg Food, water and other resources are limiting.
   - Only a fraction of the offspring in a population will
   live to produce offspring, so that the number of
   individuals in a population remains fairly constant.
3. Variation
   - Individuals in a population vary greatly in their
   characteristics.

4. Adaptation
   - Some variations enable individuals to produce more
   offspring than other individuals.

5. Natural Selection
   - Individuals having favorable traits will produce more
   offspring, and those with unfavorable traits will produce fewer
   offspring.
• Speciation
   - Given time, natural selection leads to the accumulation of
   changes that differentiate groups from one another, such that a
   new species may arise.
The Theory of Natural Selection

 The Theory of Evolution states that the present species
  have descended, with modifications, from species that
  existed in the past.

 Species are not fixed, unchanging things, but are
  constantly evolving.

 Evolution is therefore the constant change that has
 occurred in our world since its beginning to the present
 time
THE ORIGIN OF SPECIES
Variation in a population. Individuals in this
population of Asian ladybird beetles vary in color and spot pattern.
Natural selection may act on these variations only if (1) they are
heritable and (2) they affect the beetles’ ability to survive and
reproduce.
Overproduction of offspring.
A single puffball fungus can produce billions of offspring.
If all of these offspring and their descendants survived to maturity,
they would carpet the surrounding land surface.
The Theory of Natural Selection




        Darwin’s Finches
Industrial Melanism
       The Peppered Moth (Biston betularia)




Before Industrial revolution   After Industrial revolution
Evolution of drug-resistant pathogens


Evolution of insecticide resistance pests

Giraffe Necks
Hugo De Vries: Theory of Mutation (1901)

• Dutch Botanist studied
  who studied the plant,
  Evening Primrose.
• Upon observing wild
  evening primrose, Hugo
  found that they were
  different from the
  cultivated primrose
• He took some of the wild primrose seeds back to his
  garden where he made many new variations.

• He bred the plants and followed traits through several
  generations, like mendel.




               The evening primrose Plant
Hugo De Vries: Theory of Mutation (1901)

• Evolution has taken place
  through the accumulation
  of fortuitous variations or
  Mutations

• Due to the appearance of
  a series of changes in the
  germ-plasm or genes

• These      may      be
  pronounced or minor
Neo Lamarckism

This is reinvention of the Lamarck’s theory of Inheritance of
acquired characters. Where constant use tend to strengthen the
organ whereas failure of the use leads to atrophy

 Lamarck rejected the notion that the environment would
 directly affect organismal traits and neo lamarckism
 incorporates these things.

                       Neo-Darwinism
Neo-Darwinism is incorporated in the modern synthesis theory.
Darwinian evolution through Natural selection with Mendelian
genetics, the latter being a set of primary tenets specifying that
evolution involves the transmission of characteristics from parent
to child through the mechanism of genetic transfer, rather than
the 'blending process'
Modern Synthesis Theory

        Important points of Modern Evolutionary Theory
 All evolutionary phenomena can be explained in a way
consistent with known genetic mechanisms.

Evolution is gradual: small genetic changes regulated by natural
selection accumulate over long periods. Discontinuities amongst
species are explained as originating gradually through
geographical separation and extinction.

Natural selection is by far the main mechanism of change.

Thinking in terms of populations rather than individuals, is
primary: the genetic diversity existing in natural populations is a
key factor in evolution

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Evolution

  • 1. Evolution Popular Belief • Prior to the studies of Charles Darwin, the most widespread belief was that all known species were created at the same time and remained unchanged throughout history. • Some scientists at the time believed that features an individual acquired during a lifetime could be passed onto its offspring, and the species could gradually change to fit an environment better. A scientific theory is NOT just a belief, it is a well-supported tested explanation. 2.1
  • 2. Evolution Descent with Modification Evolution means change over time Evolution can also be defined more narrowly as a change in the genetic composition of a population from generation to generation.
  • 3. Fossils • A fossil is the preserved • Fossils are found in remains or imprints of sedimentary rock ancient organisms. formed when sand and silt settle to the bottom • Someone who studies of water. fossils is called a paleontologist.
  • 4. Types of radioactive dating • Carbon dating: 2 types • Potassium-Argon dating is used for rocks that are • 14C:12C suspected billions of years – Radioactive 14C old. – Non-radioactive 12C, most • In older rocks the ratio of common Argon to 40K will be higher than in younger rocks.
  • 5. Evidences for Evolution Fossil Record claws wing-like forelimbs teeth thin ribs long tail feathers Reptile-like features Replica of Archaeopteryx fossil; half bird half reptile Bird-like features
  • 6. Evidence from comparative Anatomy Vertebrate Limbs The basic pattern of the vertebrate limb is represented by the human arm skeleton shown below ball and group of hinge joint small socket joint bones single bone two bones five jointed digits
  • 7. 19 The forelimbs of the following vertebrates show the basic pattern of limb bones with modifications which are adapted to their methods of locomotion. The basic pattern suggests they have evolved from a common ancestor Lizard Bird Dolphin Bat
  • 8. DNA evidence Another important line of evidence for evolution comes from DNA analysis Organisms which have much of their DNA in common must be closely related, i.e. they have split from a common ancestor comparatively recently (in geological terms) For example, humans and chimpanzees have 99% of their DNA in common which suggests a close relationship and relatively ‘recent’ divergence from a common ancestor
  • 9. Evolutionary Theory Evolution is the process in which significant changes in the inheritable traits (genetic makeup) of a species occur over time. The validity and value of evolutionary biology are based on continual analysis and interpretation of accumulating science data. In words of Charles Darwin evolution is Descent with modification
  • 10. Theories of Organic Evolution  Historically three theories 1. Lamarck’s Theory of Evolution, 2. Darwin’s Theory of Evolution, 3. De Vries Mutation Theory of Evolution.  The views of most modern biologists combine the second and third  No evidence of the first theory, that of Lamarck’s Use and disuse theory.
  • 11. Jean Baptiste Lamarck, 1744-1829 • French geologist and naturalist • published Philosophie Zoologique in 1809 • Speculated about mechanisms of biological evolution, offered no evidence
  • 12. In 1809 Jean-Baptiste Lamarke hypothesized that “simple” organisms emerge spontaneously and evolve to greater “complexity”. Principles to his theory 1. Organisms constantly strive to improve themselves. 2. Most used body structures develop and unused waste away. (Use and Disuse) 3. A structure is modified by use or disuse, the modification is inherited to the offspring. (Inheritance of acquired characteristics).
  • 13. • Structural variations are due to functional needs – “to use or disuse” • Use of a structure increases its size. • Failure to use it decreases its size • These variations (now referred to as “acquired characteristics” are inherited. • There is no supporting evidence in modern biology to support Lamarck’s idea
  • 14.
  • 15. Disproving Lamarck • Characteristics acquired during the lifetime of a parent are not passed onto the offspring • An athlete who develops a large muscle mass through training does not have children who already possess this large muscle mass • Ernst Haeckel In an attempt to disprove Lamarckism he is said to have cut off the tails of mice for several generations • The babies born from this line of tailless mice still grew tails as long as their ancestors • This was not exactly a fair test as the mice had not stopped using their tails in an attempt to adapt to their environment •
  • 16. Charles Darwin: The Theory of Natural Selection • His theory was effectively presented to the world in 1859 when his book “The Origin of Species” was published. • Darwin’s Theory rests on 5 principles:
  • 17. 5 principles of Darwin’s Theory 1. Overproduction - All species have a tendency and the potential to increase at a geometric rate. All organisms produce more organisms but only a few survive. 2. Competition - The conditions supporting life are limited. For eg Food, water and other resources are limiting. - Only a fraction of the offspring in a population will live to produce offspring, so that the number of individuals in a population remains fairly constant.
  • 18. 3. Variation - Individuals in a population vary greatly in their characteristics. 4. Adaptation - Some variations enable individuals to produce more offspring than other individuals. 5. Natural Selection - Individuals having favorable traits will produce more offspring, and those with unfavorable traits will produce fewer offspring. • Speciation - Given time, natural selection leads to the accumulation of changes that differentiate groups from one another, such that a new species may arise.
  • 19. The Theory of Natural Selection  The Theory of Evolution states that the present species have descended, with modifications, from species that existed in the past.  Species are not fixed, unchanging things, but are constantly evolving.  Evolution is therefore the constant change that has occurred in our world since its beginning to the present time
  • 20. THE ORIGIN OF SPECIES
  • 21. Variation in a population. Individuals in this population of Asian ladybird beetles vary in color and spot pattern. Natural selection may act on these variations only if (1) they are heritable and (2) they affect the beetles’ ability to survive and reproduce.
  • 22. Overproduction of offspring. A single puffball fungus can produce billions of offspring. If all of these offspring and their descendants survived to maturity, they would carpet the surrounding land surface.
  • 23. The Theory of Natural Selection Darwin’s Finches
  • 24. Industrial Melanism The Peppered Moth (Biston betularia) Before Industrial revolution After Industrial revolution
  • 25. Evolution of drug-resistant pathogens Evolution of insecticide resistance pests Giraffe Necks
  • 26. Hugo De Vries: Theory of Mutation (1901) • Dutch Botanist studied who studied the plant, Evening Primrose. • Upon observing wild evening primrose, Hugo found that they were different from the cultivated primrose
  • 27. • He took some of the wild primrose seeds back to his garden where he made many new variations. • He bred the plants and followed traits through several generations, like mendel. The evening primrose Plant
  • 28. Hugo De Vries: Theory of Mutation (1901) • Evolution has taken place through the accumulation of fortuitous variations or Mutations • Due to the appearance of a series of changes in the germ-plasm or genes • These may be pronounced or minor
  • 29. Neo Lamarckism This is reinvention of the Lamarck’s theory of Inheritance of acquired characters. Where constant use tend to strengthen the organ whereas failure of the use leads to atrophy Lamarck rejected the notion that the environment would directly affect organismal traits and neo lamarckism incorporates these things. Neo-Darwinism Neo-Darwinism is incorporated in the modern synthesis theory. Darwinian evolution through Natural selection with Mendelian genetics, the latter being a set of primary tenets specifying that evolution involves the transmission of characteristics from parent to child through the mechanism of genetic transfer, rather than the 'blending process'
  • 30. Modern Synthesis Theory Important points of Modern Evolutionary Theory  All evolutionary phenomena can be explained in a way consistent with known genetic mechanisms. Evolution is gradual: small genetic changes regulated by natural selection accumulate over long periods. Discontinuities amongst species are explained as originating gradually through geographical separation and extinction. Natural selection is by far the main mechanism of change. Thinking in terms of populations rather than individuals, is primary: the genetic diversity existing in natural populations is a key factor in evolution