Memes are units of social information that can be transmitted from person to person through ideas or beliefs, similar to how genes are transmitted biologically. The LOLcat meme originated from image macros on forums and imageboards in the early 2000s, gaining popularity through sites like SomethingAwful, Fark, and 4chan sharing funny pictures of cats with misspelled text. By the mid-2000s, the LOLcat meme had fully emerged and was spreading widely on the internet, incorporating previous meme phrases and humor while popularizing the LOLspeak slang language.