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Homeostasis
Regulating vs Conforming
Difference between regulating and conforming
Why is regulation more energy expensive than
conforming?
Adaptation needs to outweigh the needs of the body
more than the benefits of the body
Balances
The natural environment is a system of balances
 Law of thermodynamics
Input and output should be equal
 When will there be an imbalance?
Thermoregulation
Temperature affects a lot of things
- Proteins
- Membranes
- What else?
Four physical processes of heat gain and loss
Conduction
Convection
Radiation
Evaporation
What is a wind chill effect?
Endothermy vs Ectothermy
Define endotherm/ ectotherm
True or false: ectotherms are always cold-blooded.
Advantages of endothermy
Longer vigorous activities than ectotherms
 Sustained activity is only possible in endotherms
 Thermal problems living in a terrestrial
environment is resolved through endothermy; e.g.
endotherms can live in below-freezing production-
temperature that deactivate the metabolism of
ectotherms
Disadvantage of endothermy
Thermoregulators invest more energy in their
activity than conformers, thus, increasing the energy
intake of regulators.
Question: why is ectothermy a good strategy in living
in a new environment?
Adaptation of animals that
thermoregulate
• Adjusting the rate of heat exchange between the
animal and its surroundings
• e.g. insulation such as feathers, fat
• changes in the circulatory system-
vasodilation/vasoconstriction
• countercurrent heat exchange- arteries are in
opposite direction that of veins in the extremities;
heat exchange is determined by physiology or
environment
Cooling through evaporative heat loss
Behavioural response- posture or movement
(migration/hibernation/estivation/winter sleep)
Changing the rate of metabolic heat- applies only to
endotherms
Mechanisms of temperature control
Mammals/ birds
 Constant heat loss
 Constant heat production
 Nonshivering thermogenesis vs shivering thermogenesis
Nonshivering thermogenesis (NST)
Hormones- inc metabolic activity of mitochondria
rather than production of ATP
Brown fat
Insulation
Blubber
Panting and evaporative cooling
Increase rate of heat exchange
How does fur/hair help in insulation?
Amphibians and Birds
Movement is key to heat production
Production of mucus to counter evaporative cooling
Reptiles have scales that inc skin’s SA
Vasoconstriction in extremities of marine iguana
FIshes
Mostly are conformers
Endothermic fishes use countercurrent heat
exchange system
Presence of special heat generating organs
Invertebrates
Aquatic invertebrates- mainly thermoconformers
Terrestrial invertebrates- same as vertebrate
ectotherms
Flying insects- smallest endotherms
Flying insects
Generate heat through flight muscles
Countercurrent exchange mechanisms
 Shutdown mechanisms during hot weather
Uses shivering in incubation
Huddling
Evaporative cooling
fanning
Thermostat
Controls temperature
Warm receptors- indicate inc in temp
Cold receptors- indicate dec in temp
What kind of mechanisms will each receptor
activate?
Below normal range?
Above normal range?
Acclimation vs acclimatization
Acclimation- laboratory
Acclimatization- natural environment
Antifreeze- used by ectotherms in sub-zero
environment
Proteins- e.g. Heat shock proteins
Torpor
Physiological state of low activity and low
metabolism
Hibernation vs winter sleep
Estivation
Daily torpor
Osmoregulation
Functions in maintaining the composition of the
cell’s cytoplasm
Mostly done indirectly
Open circulatory- uses hemolymph
Close circulatory- use interstitial fluid
Kidneys are specialized organs in maintaining the
composition of the body’s fluid composition
Transport epithelia
e.g. transport epithelium face the outside
environment to release unwanted solutes but have
tight junction in between cells to inhibit back flow;
functions like the Casparian strip of plants
Ammonia
Most common in aquatic animals
Can easily pass through membranes via diffusion
Invertebrates release ammonia all throughout the
body
Fish release ammonia in the form of ammonium ions
through the gills (kidneys excrete only minimal
amount)
Freshwater fishes excrete NH4 ions but also take in
Na ions through the gill epithelium to have a higher
concentration of Na ions compared to the
environment
Urea
less toxic compared to ammonia
Need less water in eliminating
Used by mammals, adult amphibians, marine fishes
and turtles
Ammonia+CO2
Transported via the circulatory system and filtered
in the kidneys
Can be transported in high concentration due to low
toxicity
Uses more energy
Animal adaptation: amphibians in water excrete
ammonia but excrete urea in land, what is the
advantage of this lifestyle?
Uric Acid
relatively nontoxic nitrogenous waste
Insoluble in water and excreted as semisolid paste
Advantage: low water loss
Disadvantage: highly expensive
Present in land snails, insects, birds, reptiles
Osmoconformers vs osmoregulators
Osmoconformers- animals that have the same
concentration of body fluid and of the external
environment; live in relatively stable environment
Osmoregulators- maintains the concentration of
body fluid; body fluid is not isoosmotic with that of
the environment
Stenohaline- animals that cannot tolerate broad
change in solute concentration
Euryhaline- animals that can tolerate substantial
change in external osmolarity, e.g. salmon
Homeostasis

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Homeostasis

  • 2. Regulating vs Conforming Difference between regulating and conforming Why is regulation more energy expensive than conforming?
  • 3. Adaptation needs to outweigh the needs of the body more than the benefits of the body
  • 4. Balances The natural environment is a system of balances  Law of thermodynamics Input and output should be equal  When will there be an imbalance?
  • 5. Thermoregulation Temperature affects a lot of things - Proteins - Membranes - What else?
  • 6. Four physical processes of heat gain and loss Conduction Convection Radiation Evaporation
  • 7. What is a wind chill effect?
  • 8. Endothermy vs Ectothermy Define endotherm/ ectotherm True or false: ectotherms are always cold-blooded.
  • 9. Advantages of endothermy Longer vigorous activities than ectotherms  Sustained activity is only possible in endotherms  Thermal problems living in a terrestrial environment is resolved through endothermy; e.g. endotherms can live in below-freezing production- temperature that deactivate the metabolism of ectotherms
  • 10. Disadvantage of endothermy Thermoregulators invest more energy in their activity than conformers, thus, increasing the energy intake of regulators.
  • 11. Question: why is ectothermy a good strategy in living in a new environment?
  • 12. Adaptation of animals that thermoregulate • Adjusting the rate of heat exchange between the animal and its surroundings • e.g. insulation such as feathers, fat • changes in the circulatory system- vasodilation/vasoconstriction • countercurrent heat exchange- arteries are in opposite direction that of veins in the extremities; heat exchange is determined by physiology or environment
  • 13. Cooling through evaporative heat loss Behavioural response- posture or movement (migration/hibernation/estivation/winter sleep) Changing the rate of metabolic heat- applies only to endotherms
  • 14. Mechanisms of temperature control Mammals/ birds  Constant heat loss  Constant heat production  Nonshivering thermogenesis vs shivering thermogenesis
  • 15. Nonshivering thermogenesis (NST) Hormones- inc metabolic activity of mitochondria rather than production of ATP Brown fat Insulation Blubber
  • 16. Panting and evaporative cooling Increase rate of heat exchange
  • 17. How does fur/hair help in insulation?
  • 18. Amphibians and Birds Movement is key to heat production Production of mucus to counter evaporative cooling Reptiles have scales that inc skin’s SA Vasoconstriction in extremities of marine iguana
  • 19. FIshes Mostly are conformers Endothermic fishes use countercurrent heat exchange system Presence of special heat generating organs
  • 20. Invertebrates Aquatic invertebrates- mainly thermoconformers Terrestrial invertebrates- same as vertebrate ectotherms Flying insects- smallest endotherms
  • 21. Flying insects Generate heat through flight muscles Countercurrent exchange mechanisms  Shutdown mechanisms during hot weather Uses shivering in incubation Huddling Evaporative cooling fanning
  • 22. Thermostat Controls temperature Warm receptors- indicate inc in temp Cold receptors- indicate dec in temp What kind of mechanisms will each receptor activate?
  • 24. Acclimation vs acclimatization Acclimation- laboratory Acclimatization- natural environment
  • 25. Antifreeze- used by ectotherms in sub-zero environment Proteins- e.g. Heat shock proteins
  • 26. Torpor Physiological state of low activity and low metabolism Hibernation vs winter sleep Estivation Daily torpor
  • 27. Osmoregulation Functions in maintaining the composition of the cell’s cytoplasm Mostly done indirectly Open circulatory- uses hemolymph Close circulatory- use interstitial fluid Kidneys are specialized organs in maintaining the composition of the body’s fluid composition
  • 28. Transport epithelia e.g. transport epithelium face the outside environment to release unwanted solutes but have tight junction in between cells to inhibit back flow; functions like the Casparian strip of plants
  • 29. Ammonia Most common in aquatic animals Can easily pass through membranes via diffusion Invertebrates release ammonia all throughout the body Fish release ammonia in the form of ammonium ions through the gills (kidneys excrete only minimal amount) Freshwater fishes excrete NH4 ions but also take in Na ions through the gill epithelium to have a higher concentration of Na ions compared to the environment
  • 30. Urea less toxic compared to ammonia Need less water in eliminating Used by mammals, adult amphibians, marine fishes and turtles Ammonia+CO2 Transported via the circulatory system and filtered in the kidneys
  • 31. Can be transported in high concentration due to low toxicity Uses more energy Animal adaptation: amphibians in water excrete ammonia but excrete urea in land, what is the advantage of this lifestyle?
  • 32. Uric Acid relatively nontoxic nitrogenous waste Insoluble in water and excreted as semisolid paste Advantage: low water loss Disadvantage: highly expensive Present in land snails, insects, birds, reptiles
  • 33. Osmoconformers vs osmoregulators Osmoconformers- animals that have the same concentration of body fluid and of the external environment; live in relatively stable environment Osmoregulators- maintains the concentration of body fluid; body fluid is not isoosmotic with that of the environment
  • 34. Stenohaline- animals that cannot tolerate broad change in solute concentration Euryhaline- animals that can tolerate substantial change in external osmolarity, e.g. salmon